Plants terrestrial or epilithic, evergreen or seasonally green, creeping, (5-)15-45 cm or more. Rhizophores at intervals. Main stems pinnately branched, stramineous; primary leafy branches 3-9 pairs or more, once or twice pinnately branched, secondary branches once or twice forked, branchlets sparse, adjacent primary branches on main stem 1-2(-5) cm apart, leafy portion of main stem including leaves 4.5-6(-8.5) mm wide at middle, ultimate branches 3.6-4.2(-6.5) mm wide including leaves. Axillary leaves on branches symmetrical or asymmetrical, ovate-lanceolate, (1.2-)2-3(-3.8) × (0.6-)1.2-1.8 mm, margin subentire, denticulate or ciliolate in basal half. Dorsal leaves asymmetrical, obliquely ovate, 1.6-2(-2.8) × 0.5-0.8(-1.2) mm, base subcordate, margin denticulate or shortly ciliolate (at least at base), apex aristate, arista short to more than 1/2 length of dorsal leaf and often reflexed. Ventral leaves spreading or slightly deflexed, asymmetrical, oblong-ovate, (1.6-)2-3(-3.6) × 1-1.8 mm, apex acute or obtuse; basiscopic margin subentire or minutely denticulate; acroscopic base enlarged, broader, overlapping stem and branches, margin subentire or minutely denticulate. Strobili solitary, terminal, compact, tetragonal, 3-17 × 2-4.6 mm; sporophylls uniform, white-margined, ovate, sharply carinate, margin denticulate or ciliolate, apex aristate; megasporophyll only 1 at base on lower side of strobilus, elsewhere microsporophylls or megasporophylls in basal portion on lower side, or megasporophylls and microsporophylls at intervals; microsporangia elliptic; microspores yellowish orange, megaspores whitish.
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