Plants terrestrial, evergreen, suberect or ascending from decumbent base, 25-45 cm, without creeping rhizomes or stolons. Rhizophores branched from base to middle of main stem. Main stems branched from lower part upward, pinnately branched, stramineous, 1-3 mm in diam. in lower part, oval or subquadrangular, glabrous; primary leafy branches 3-6 pairs, 2 or 3 times pinnately branched, secondary branches once pinnately branched, tertiary branches forked, branchlets sparse, adjacent primary branches on main stem 3-6 cm apart, leafy portion of main stem including leaves 0.7-1 mm wide at middle, ultimate branches 4-7 mm wide including leaves. Axillary leaves on branches symmetrical, narrowly ovate to triangular, 1.8-3 × 0.9-1.4 mm, base exauriculate, margin denticulate. Dorsal leaves on branches imbricate, oblong-ovate, ovate-elliptic, or narrowly ovate, 1.1-2.7 × 0.4-1.4 mm, carinate or strongly carinate, base cuneate or obliquely subcordate, margin denticulate, apex acuminate to aristate, parallel to axis. Ventral leaves on branches contiguous or overlapping, slightly ascending, oblong-falcate, 2.3-4.4 × 1-1.8 mm, apex obtuse, subacute, or apiculate; basiscopic base slightly dilated, margin subentire, denticulate at base; acroscopic base enlarged, broader, overlapping stem and branches, margin denticulate. Strobili solitary or in pairs, terminal, compact, tetragonal, 5-30 × 1-2 mm; sporophylls uniform, not obviously white-margined, ovate-triangular, carinate, margin denticulate, apex acuminate; megasporophylls and microsporophylls at intervals, or megasporophylls in basal portion on lower side of strobilus; microsporangia elliptic-oblong, relatively thick, marginal cells differentiated, longer; microspores yellowish orange, megaspores whitish, without equatorial flange.