Plants terrestrial or epilithic, evergreen or seasonally green, erect or ascending from decumbent base, (5-)15-20(-30) cm, with creeping subterranean rhizome and stolons. Rhizophores restricted to base of stem or borne on creeping rhizomes and stolons. Main stems branched from middle or lower part upward, pinnately branched, stramineous or reddish (when alive), 0.4-1.4 mm in diam. in lower part, terete, sulcate; primary leafy branches 3-5 pairs, 2 or 3 times pinnately branched, secondary branches once or twice pinnately branched, tertiary branches forked or simple, branchlets sparse or dense, adjacent primary branches on main stem 1-5 cm apart, ultimate branches (2.2-) 3-3.5(-5.5) mm wide including leaves. Axillary leaves on main stems larger than those on branches, ovate, base not peltate, truncate; axillary leaves on branches asymmetrical, ovate-lanceolate, (1.4-)2-2.4(-2.9) × (0.5-)0.8-1(-1.3) mm, base exauriculate, margin denticulate or ciliolate. Dorsal leaves ± symmetrical, those on main stems obviously larger than those on branches; dorsal leaves on branches approximate, ovate or ovate-lanceolate, 0.9-2 × 0.3-0.8 mm, carinate or not carinate, base subcordate, not peltate, margin denticulate or ciliolate (at base), apex aristate, often reflexed or arista curved. Ventral leaves asymmetrical, those on main stem obviously larger than those on branches; ventral leaves on branches distant, slightly ascending, ovate-lanceolate, narrowly ovate, or triangular, 1.7-3.2 × 0.6-1.2 mm, apex acute; basiscopic base rounded, denticulate or ciliolate at base (denticulate to apex); acroscopic base enlarged, broader, overlapping stem and branches, margin shortly ciliolate at base, denticulate to apex. Strobili solitary, terminal, compact, dorsiventrally complanate or subcomplanate, 5-18 × 1.3-3 mm; sporophylls unlike sterile leaves, slightly or strongly dimorphic, resupinate, white-margined; dorsal sporophylls ovate-lanceolate, margin ciliolate or denticulate, apex acuminate, with sporophyll-pteryx incomplete and ciliolate or denticulate; ventral sporophylls ovate, carinate, margin denticulate or ciliolate, apex aristate or acuminate; megasporophylls and microsporophylls at intervals or megasporophylls in basal or upper portion on lower side of strobilus; microsporangia orbicular, rather thin, cells uniform; microspores orange-red or red, megaspores pale yellow or yellowish orange.
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