Plants on rock or terrestrial, forming loose or compact mats. Stems radially symmetric, creeping or decumbent, not readily fragmenting, irregularly forked, without budlike arrested branches, tips straight; main stem long, indeterminate, lateral branches determinate, ascending, 1--2-forked. Rhizophores borne on upperside of stems, throughout stem length, 0.23--0.36(--0.4) mm diam. Leaves monomorphic, in ± alternate pseudowhorls of 4, tightly or loosely appressed, ascending, green, linear-lanceolate, (1.5--)1.8--3.5 X 0.39--0.66 mm; abaxial ridges well defined; base rounded and adnate or cuneate and slightly decurrent on fleshy, loosely appressed stem leaves (from wet places), pubescent, seldom glabrous; margins short-ciliate to denticulate, cilia transparent, spreading at base, dentiform, and ascending toward apex, 0.03--0.06(--0.1) mm; apex keeled and obtuse, sometimes attenuate or plane and attenuate, abruptly short-to long-bristled; bristle transparent to whitish, puberulent, sometimes breaking off, (0.16--)0.2--0.46(--0.9) mm. Strobili often paired, 1--4.5(--9) cm; sporophylls deltate-ovate (mostly on exposed and compact mats) or lanceolate-ovate (on loose, spreading mats from wet places), abaxial ridges well defined, base glabrous, margins short-ciliate to denticulate, apex keeled, abruptly short-bristled, seldom tapering into bristle.