Plants in small to large mats, usually golden to yellow-green, glossy or dull. Stems reddish green or sometimes red-brown, reclining or sometimes erect or arching, complanate-foliate or not, irregularly branched to pinnate; pseudoparaphyllia foliose or filamentous. Stem leaves erect-spreading, wide-spreading, falcate-secund, or homomallous, lanceolate to broadly ovate, tapering gradually or abruptly to apex; margins often toothed in acumen; apex acute to acuminate; ecostate or costa double, short; alar region well defined, pigmented, cells in proximal row strongly elongate and inflated, or not inflated and walls thicker; laminal cells usually flexuose, smooth or papillose (serially or 1-papillose); medial cells occasionally linear. Branch leaves smaller, narrower; alar cells less strongly differentiated. Specialized asexual reproduction by flagelliferous branches or axillary filamentous propagula. Sexual condition synoicous, dioicous, phyllodioicous, or autoicous. Seta reddish brown, red, or light brown, elongate, smooth. Capsule inclined or erect, cylindric or ovoid; exothecial cell walls usually collenchymatous, irregularly thickened; operculum usually rostrate; peristome double. Calyptra cucullate [mitrate].