Annual or perennial herbs, shrubs, trees, or climbers, not usually strongly scented, lacking latex. Lvs usually alternate, rarely opposite, whorled or all basal. Involucral bracts usually in 1-2 rows, rarely more, usually foliaceous, sometimes subtended at base by small supplementary bracts. Receptacle without scales. Capitula usually ☿, rarely unisexual, either heteromorphic with 1-(several) rows of ligulate ray florets and actinomorphic disc florets, or homomorphic-discoid. Anthers rounded, acute or sagittate at base. Style branches usually each with 2 stigmatic lines, sometimes lines fused. Achenes usually homomorphic, rarely heteromorphic, often with c. 10 ribs, terete or compressed; pappus of 1-few rows of smooth or toothed hairs, sometimes caducous or 0.