Dioecious tendrilled lianas, densely rufous-pubescent except sparsely so on the upper surface of the leaflets; stems 5-ribbed; wood simple or composite with at least some of the ribs consisting of separate bundles. Leaves biternate; petioles mostly 3-6.5 cm long, ribbed above; rachis margined; leaflets elliptic, blunt to acuminate, abruptly tapered to the base, especially the lateral leaflets, remotely ser-rate-dentate, the terminal leaflets mostly 7-13 cm long, 3.5-5.5 cm wide, the other leaflets gradually smaller. Thyrses paniculate, terminal or in the upper leaf axils, the branches to 15 cm long; pedicels to 1 cm long. Flowers white, ca. 7 mm long, unisexual; sepals 5, tomentulose; petals 4, oblong-obovate, ciliolate, the scale of the anterior petals to 2/3 the length of the petal, its crest yellow, thin, emarginate, the appendage villous, slender, pendent to just above glands (scales of lateral petals lacking appendages); rounded disc glands subtending both anterior petals, minutely pubescent on the lower margins, depressed on the outer face, white in staminate flowers, purple at the apex in pistillate flowers; pistillate flowers with stamens to 3.5 mm long, the style 3-4 mm long, the stigmas 3, ca. 1 mm long; staminate flowers with the 5 longer stamens to 6.5 mm long, the shorter ones to 4 mm long, the filaments sparsely villous, the pistil nonfunctional. Fruits sub-rectangular-oblong, ca. 4 cm long, the seed-bearing part rufous-pubescent, sub-apical, hirtellous and setose with a hornlike projection, the wings scarcely if at all constricted below the seed-bearing part, hirtellous.
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