Shorea macrophylla (De Vriese) P.S.Ashton

Medium-sized to large tree. Young twig, leaf bud, stipules, petiole, leaf beneath and midrib above persistently densely evenly caducous pale brown pubescent. Twig 2.5-4 by 2-3 mm ø apically, compressed, becoming smooth, glabrous; stipule scars to 1 mm thick, amplexicaul, pale cream, +-horizontal, smooth, glabrous. Bud 12-18 by 4-6 mm, compressed, hastate, narrowly obtuse. Stipule to 5 by 1.3 cm, broadly hastate, obtuse, constricted at base. Leaves 17-35 by 10-14 cm, elliptic-oblong, +-coriaceous; base obtuse or subcordate; acumen to 1.5 cm long; nerves 11-20 pairs, prominent beneath, well spaced, at 55°-65°; tertiary nerves distinct, well spaced, scalariform, at 90° to nerves; midrib prominent, rounded, beneath, broad and applanate above; petiole 1.5-3 cm long. Panicle to 17 cm long, terminal or axillary, ribbed and compressed when dry, shortly evenly persistently buff pubescent, singly branched; bracteoles to 1.2 by 0.5 cm, oblong, subacute. Flower bud to 8 by 5 mm, broadly or narrowly ellipsoid. Calyx lobes densely pale brown pubescent outside, glabrous within; lobes subequal, broadly elliptic to deltoid, obtuse to acuminate. Petals pale pink, broadly ovate, obtuse, densely pubescent on parts exposed in bud. Stamens 15, of 2 lengths, the inner 5 an anther's length longer than the outer 10; filaments broad, compressed, united in a tube round the ovary, tapering distally and filiform below the oblong anthers; appendage to connective c. 2 x length of anther, slender, glabrous. Ovary ovoid, densely pubescent in the distal half; stylopodium and style glabrous, spindle shaped, as long as ovary. Fruit calyx glabrous; 3 longer lobes to 11 by 3 cm, coriaceous, oblong, obtuse, to 1.5 cm broad above the to 1.8 by 2.3 cm horizontally elliptic deeply saccate thickened base; 2 shorter lobes to 8 by 1.5 cm, similar at base, base obscured by larger lobes. Nut to 6 by 4 cm, obovoid, persistently evenly shortly pale buff pubescent; style remnant, small, short, acute.

A canopy tree, locally abundant in mixed dipterocarp forest, growing on clay-rich periodically flooded alluvium and river banks, uncommon on hillsides, at elevations below 600 metres.

Uses. The principal source of the Bornean Illipe nut; sometimes planted.

Can be grown by seedlings. Seeds needs soaking.