Shorea quadrinervis Slooten

Medium-sized, occasionally large, tree. Young parts shortly densely tawny pubescent, largely caducous on stipule, leaf bud and leaf beneath; leaf above glabrous but for midrib. Twig c. 3 mm ø at apex, terete, stout, strongly ridged below stipule scars, becoming minutely cracked longitudinally; stipule scars c. 3 mm long, c. 1 mm broad, pale, falcate or amplexicaul-descending. Bud to 2 by 1.3 cm, loosely enclosed by stipular bud scales, compressed, subcordate. Stipule to 2.5 by 1.3 cm, ovate to subcordate, subpersistent, glabrescent, persistently ridged along the 5 nerves; stipule pairs sometimes united at base. Leaves 10-18 by 5-8 cm, broadly ovate to elliptic, strongly cupped; margin usually slightly revolute; base obtuse; acumen to 1.3 cm long; nerves c. 4 pairs, strongly curved, at c. 40°-70° with 1 or more secondary nerves of varying length; tertiary nerves slender, scalariform, at c. 90° to nerves; midrib depressed above; petiole 8-10 mm long. Panicle to 28 cm long, terminal or axillary, large, spreading, lax, terete or angular, densely persistently pale rust pubescent; doubly, sometimes trebly branched, branchlets 1-8 cm long, straight, bearing to 6 secund flowers; bracts as the stipules, +-caducous; bracteoles to 2 by 1 mm, ovate to elliptic, obtuse, densely grey pubescent outside, puberulent within. Flower bud to 4.5 by 5.5 mm, broadly elliptic to subglobose, obtuse. Calyx shortly pale grey pubescent outside, glabrous within; lobes +-patent, subequal, ovate, the inner 2 slightly more acute. Petals pink, paler at margins, elliptic, obtuse, densely pubescent on parts exposed in bud. Stamens 15, the inner 5 longest, the outer 10 sometimes vestigial; filaments tapering gradually; anther subglobose; appendage to connective shorter than anther, becoming reflexed. Ovary and stylopodium conical, densely pubescent; style short, frequently expanded at the stigma. Fruit calyx shortly sparsely pubescent; 3 longer lobes to 8 by 1.3 cm, narrowly spatulate, obtuse, 2-3 mm broad above the c. 5 by 4 mm thickened base; 2 shorter lobes to 5.5 by 0.7 cm, unequal, tapering to c. 2 mm broad, base as in longer lobes. Nut to 1.5 by 0.7 cm, narrowly ovoid, shortly buff pubescent; style remnant c. 1.5 mm long, slender, acute.

A canopy to emergent tree in mixed dipterocarp forest, common on leached, yellow, sandy clay soils on low hills and on skeletal soils on ridges; occasionally to elevations of 700 metres.

Can be grown by seedlings. Seeds needs soaking.