Plants perennial; taproot stout; caudex branched, woody. Stems several, erect, freely branched, leafy, elongate, 30-80 cm, scabrid-puberulent and glandular-viscid. Leaves 2 per node; basal long-petiolate, blade oblanceolate, (2-)5-18 cm × 5-30 mm, base narrowed into petiole, apex ± acute, glandular-puberulent on both surfaces; cauline not greatly reduced in distal stem, blade 5-15 cm × 10-20 mm, viscid glandular-pubescent, sparsely so in shade forms, proximal petiolate and blade oblanceolate, distal sessile and blade lanceolate, apex acute. Inflorescences open, elongate, dichotomously branched, branches ascending, bracteate; bracts leaflike, reduced distally. Pedicels ascending, may be partially deflexed, rather slender, longer than calyx, viscid glandular-pubescent with septate hairs, septa colorless. Flowers: calyx prominently 10-veined, tubular in flower, 8-12 × 2-4 mm, swelling to campanulate and 5-7 mm broad in fruit, not contracted proximally around carpophore, viscid glandular-pubescent, especially on veins, veins parallel, green, with pale commissures, those to lobes broadened distally, lobes erect, narrowly lanceolate, 3-4 mm, rigid, setose-scabrous, glandular-viscid, apex recurved; corolla greenish white, claw equaling calyx, limb 2-lobed, ca. 3 mm, appendages ca. 0.5 mm, margins erose; stamens equaling petals; styles 3, equaling petals. Capsules slightly exserted from calyx, narrowly ovoid, opening by 6 teeth; carpophore 1-2 mm. Seeds almost black, ellipsoid-reniform, ca. 1 mm, coarsely papillate; papillae inflated. 2n = 48.
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