Herbs perennial, usually rhizomatous. Stems erect or prostrate. Leaves alternate. Basal leaves long petiolate, ovate-cordate; stem leaves smaller, short petiolate. Cymes terminal, racemose or paniculate. Calyx 5-parted to near base, enlarged, becoming saccate in fruit. Corolla tubular to funnelform; throat appendages 5, shallowly 2-cleft; limb 5-parted, spreading. Stamens inserted above middle of corolla tube or at throat between appendages, exserted or included; filaments filiform. Ovary 4-divided. Style elongated; stigma capitate; ovules anatropous. Gynobase low fastigiate. Nutlets tetrahedral; margin abaxially exserted, becoming a cupular emergence; attachment scar slightly below middle adaxially.