Liana or scandent shrub, sometimes epiphytic, the stems with numerous long aerial roots, the branchlets yellowish or grayish-white to yellowish-brown or dark gray, the older branches with dark silvery gray bark. Leaves variable in shape and size, often asymmetric, chartaceous when dried; petiole 5-12(-20) mm long and 1.5-2.5 mm wide, canaliculate above; blade elliptic-oblong, oblong, obo-vate-oblong, or rarely lanceolate or obovate, (1.5-)2(-3) times as long as wide, 8-14(-16) cm long and 3.5-6(-8) cm wide, subacute, obtuse, or rounded, or sometimes acute or subacuminate apically, acute to obtuse or sometimes rounded basally; the midrib flat to slightly prominent and often sulcate above, prominent beneath, the lateral veins mostly prominent on both sides of the blade; hypophyl-lous glands variable in number and size, from small (0.5 mm) and inconspicuous to large (ca. 2 mm) and conspicuous, often 4-15 in a more or less distinct and regular row 5-12 mm from the margin and often another row of smaller glands near the margin, especially in the apical part of the blade, rarely com-pletely or almost wanting. Inflorescences lax, mostly with 15-30 flowers, the rachis 10-30 cm long, yellowish-brown puberulent, often rugose when dried; nectaries auriculate spurs, straddling the pedicels, 1 5-33 mm long, the spur clavate, 7-15 mm long, the auricles 10-20 mm long and 2-5 mm wide. Flowers fragrant, the pedicels slender to stout, especially in fruit, from 1 cm at the apex of the inflorescence to 2-4(-5.5) cm long at the base, 1-2 mm thick, puberulent, often dark and finely wrinkled when dried; bracteoles semi-to suborbicular, 2-2.5 long and ca. 3 mm wide; sepals suborbicular, 2-3 mm long and 3-4.5 mm wide; corolla gamopetalous, the petals mostly connate for more than half their length, 7-11 mm long, at anthesis the tube ruptured and the lobes reflexed; stamens 5, the filaments flattened, 4-6 mm long, 2-3 mm wide, connate basally, narrowing apically, the anthers ovate, 1.5-2.5 mm long and 1-1.5 mm wide; ovary 5-loculed, pentagonal, ca. 2.5 mm long and 2-3 mm in diameter, the stigma large, 5-radiate. Fruit globose, hard and woody when dried, ca. 2 cm in diameter, apiculate.