Plants moderate-sized to robust, moderately stiff-stemmed, ± lax, but not compact; green to pale yellow to yellowish brown; capitulum typically strongly convex in open grown forms, but flat and ± 5-radiate in shade forms. Stem pale green to yellowish; superficial cortex of 2 layers of enlarged, thin-walled and well differentiated cells. Stem leaves triangular, triangular-lingulate to lingulate, more than 0.8 mm, appressed, apex obtuse to broadly obtuse, erose to fimbriate; hyaline cells efibrillose and nonseptate. Branches straight and often tapering, often 5-ranked, leaves not much elongate at distal end of branches. Branch fascicles with 2 spreading and 2 pendent branches. Branch stems green, cortex enlarged with conspicuous retort cells. Branch leaves ovate-lanceolate, 1.4-2 mm, straight; slightly undulate and sharply recurved; margins entire; hyaline cells on convex surface with 1 pore per cell at cell apex, concave surface with round wall thinnings in the cell ends and angles; chlorophyllous cells triangular in transverse section and well-enclosed on concave surface. Sexual condition dioicous. Spores 22-28 µm; papillose on both surfaces; proximal laesura more than 0.5 spore radius.
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