Plants small to moderate-sized, slender and stiff, capitulum ± rounded, rarely flat-topped or stellate; green to variegated red-green especially in new growth, capitulum, and antheridial branches, without metallic sheen when dry. Stems green to reddish, superficial cortical cells aporose. Stem leaves broad-triangular to triangular lingulate, 0.9-1.2 mm, apex acute to slightly rounded, border strongly broadened at base (more than 0.3 width); hyaline cells mostly 0-1-septate, S-shaped to rhomboid. Branches not 5-ranked. Branch fascicles with 2 spreading and 2 pendent branches (rarely 1). Branch leaves 0.9-1.2 mm, ovate-lanceolate, 0.9-1.2 mm, concave, straight, apex involute; hyaline cells on convex surface with numerous round to elliptic pores along the commissures (4-8), grading from small pores near apex to large pores at leaf base, concave surface with large round pores in proximal portions of leaf. Sexual condition dioicous. Spores 19-29 µm, finely papillose on proximal surface, more coarsely papillose on distal surface, conspicuous bifurcated Y-mark sculpture on distal surface; proximal laesura less than 0.5 spore radius.