Plants medium-sized to large, pendent, green to yellow-green turning blackish brown to black on older stems and branches; corticolous or saxicolous. Primary stems creeping; secondary stems pendent, julaceous, to 200 mm long, branching regular, in widely spaced groups of 2-5 branches; in section round, central strand variable, small or absent, inner cortical cells hyaline, thin-walled, in 6 rows, outer cortical cells yellow, thick-walled in 4 or 5 rows. Leaves ± crowded and concave, ± erect wet, appressed dry; stem leaves ovate-lanceolate to oblong, 3.5 mm long; abruptly setaceous; subcordate and decurrent at base; margins plane, entire, cells not differentiated; branch leaves broadly ovate to elliptical or oblong; abruptly cuspidate to setaceous; subcordate and decurrent at base; margins plane, entire. Costa variable, mostly single, extending to upper leaf, on stem leaves extending to midleaf, percurrent or short and double on branch leaves; ventral and dorsal surface smooth; in section flat, guide cells not differentiated, ventral and dorsal surface cells thick-walled. Upper laminal cells linear-fusiform, walls weakly thickened, homogeneous, smooth; basal cells rectangular, hyaline, walls thickened and pitted, smooth; alar cells strongly differentiated, quadrate, yellowish, walls thickened. Dioicous. Perigonia on stems and branches, gemmate. Perichaetia along stem; perichaetial leaves oblong-acuminate, base sheathing, 4 mm long, leaf cells linear, fusiform. Seta 0.5-1.0 mm long, brown, smooth. Capsule immersed, ovoid, 2 mm long, smooth, black-green, neck not differentiated; exothecial cells subquadrate broadly rectangular, walls thickened, cells at mouth quadrate, neck cells smaller; stomata phaneropore. Peristome double, yellow-brown; exostome teeth long-linear, smooth, papillose, 0.7 mm high; endostome segments linear above low basal membrane, perforated, shorter than teeth, smooth; cilia absent. Operculum ± convex, 0.7 mm long. Calyptra cucullate, 1 mm long, pilose. Spores rounded, 25-35 µm, granulate, yellow.
Home