Trees or shrubs, resting buds with cataphylls. Leaves pubescent at first, sometimes becoming glabrous, entire or dentate, principal secondary veins arched and meeting within the margin. Monoecious or dioecious. Inflorescences later-al, cymose or fasciculate. Male flowers globose, turbinate or patelliform, with 4 small tepals, stamens (3-)4(-5), or rarely with a second (inner) whorl of 1-4 smaller stamens; anthers opening by a single horizontal slit. Female flowers similar to male but larger and with a smaller ostiole surrounded by 4 tepals, the upper half abscissing as a calyptra after anthesis to reveal numerous carpels; ovary pubescent; style glabrous awl-shaped. Drupes sessile or long-stipitate.