Terricolous, on damp soil; thallus, mostly erect, dendroid shoots in loose mats or tufts, green to olive-green, medium-sized; proximal branches, after initial dichotomy, single, ± 4 mm long, then a further 2 or 3 times dichotomously furcate, with 7 or 8(-12) terminal branches, linear to narrowly ovate, up to 11 mm long, 1.6-2.0 mm wide, 125 µm thick over ventrally slightly bulging costa, lacking rhizoids, but with central, brown conducting strand clearly visible and forking at dichotomies; apex rarely progressively narrowed with a reversion to stipe-like condition and arching down, mostly entire or slightly to deeply notched, bearing 2-celled slime papillae, margins of wings dentate, lacking slime papillae, plane, not undulating or hardly so, bilaterally expanded from ascending wingless stipe, 4-18 mm long, in transverse section 275 x 500 µm, arising from horizontally creeping and much branched, cylindrical brown rhizome, with numerous smooth ventral rhizoids, ± 12.5 µm wide. Wings generally unistratose, but medianly bistratose and grading into flattish costa, laterally with marginal teeth, remote or closer together, occasionally very blunt, usually with 2(or 3) forwardly directed cells, diverging at an angle of ± 45°, basal cell 62.5 x 37.5 µm (rarely with 2 adjacent cells), top cell bluntly conical, 50 x 30 µm; marginal cells ± rectangular to polygonal (40.0-)67.5-90.0 x 27.5-37.5 µm, 30-35 µm thick in transverse section, their walls and those of 1(or 2) rows of inframarginal cells generally somewhat thicker , coloured pinkish; laminal cells arching toward margins, polygonal, up to 82.5 x 55.0 µm, epidermal cells along costa narrowly rectangular or long-hexagonal, 75.0-125.0 x 37.5 µm, occasionally bearing 2-celled slime grading into flattish costa, laterally with marginal teeth, remote or closer together, occasionally very blunt, usually with 2(or 3) forwardly directed cells, diverging at an angle of ± 45°, basal cell 62.5 x 37.5 µm (rarely with 2 adjacent cells), top cell bluntly conical, 50 x 30 µm; marginal cells ± rectangular to polygonal (40.0-)67.5-90.0 x 27.5-37.5 µm. 30-35 µm thick in transverse section, their walls and those of 1(or 2) rows of inframarginal cells generally somewhat thicker , coloured pinkish; laminal cells arching toward margins, polygonal, up to 82.5 x 55.0 µm epidermal cells along costa narrowly rectangular or long-hexagonal, 75.0-125.0 x 37.5 µm, occasionally bearing 2-celled slime papillae above; chloroplasts numerous, ± 5 µm wide, entirely filling cells, or clustered along cell walls, several oil bodies also present, spindle-shaped and ± 7.5 µm long, or round when viewed end-on. Costa with central conducting strand, 37.5 x 50.0 µµm, consisting mostly of 12 small, ± 10 x 10 µm, brown, thick-walled, angular cells, surrounded above and below by 3 rows of larger, thin-walled parenchymatous cells 20.0-22.5 x 25.0-37.5 µm. Dioicous. Androecia in 1 or 2 dorsal rows over the costa; antheridia 200 x 175 µm, short-stalked. Each one individually covered and well hidden by irregularly shaped, forwardly directed scale-like involucre, 175 µm wide at base, up to 450 µm long, irregularly incised at apex. Gynoecia generally 2 per frond, dorsally situated above costa at bifurcation of conducting strand, containing several archegonia and subtended by posteriorly inserted involucre, 675 µm wide at base, deeply laciniate to filiform above, cells short-to long-rectangular, 45.0-125.0 x 27.5-47.5 µm. Calyptra thickening and enlarging into a fleshy shoot calyptra, ± 4 mm long and up to 8 cell rows or 250 um thick in transverse section, with several unfertilized archegonia remaining attached near the top. Capsule cylindrical, 1800 x 850 m, opening along several longitudinal valves, remaining attached above, wall brown, 2 cell layers thick, outer cells elongate, 62.5-107.5 x 17.5 um, walls thickened, inner cells thin-walled. Seta erect when young, 480 µm in diameter, with ± 36 cortical cells, ± 50.0 x 32.5 µm, medullary cells slightly larger, 62.5 x 37.5 µm, angular, thin-walled. Spores light brown, ± globular, 20-25 µm in diameter, ornamentation nodular, with some very irregular, broad flattened granular ridges, meshes sometimes distinct, but mostly not; proximal face with small round area with compact, punctate ornamentation. Elaters brown, hardly tapering toward ends, 305-330 x 7.5 µm, 2-spiral.
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