Monoecious trees. Branchlets lenticellate, hairy with simple hairs. Leaves paripinnate; leaflets alternate or sub-opposite, coriaceous. Inflorescence axillary, spike-like or panicle-like; cymules clustered. Flowers regular. Calyx cupular, deeply 5-lobed; lobes ovate, slightly imbricate at base. Petals 5, ± rhomboidal, longer than calyx, with 2 large, usually crested, villous scales. Disc fleshy, annular, glabrous. Stamens usually 8; filaments filiform, villous. Ovary trigonous, 3-locular; ovule 1 per locule. Fruit obovoid or subglobose, keeled at sutures, loculicidally 3-valved; valves fleshy, drying thinly coriaceous, glabrous outside, villous inside. Seeds obovoid, slightly dorsally compressed, reddish brown; aril mantle-like or oblique, bilobed, adhering to base of seed.