Plants terrestrial. Rhizome creeping, apex covered with dark, rigid bristles; bristles with a single row of cells at apex and a few rows of cells at base. Fronds monomorphic or dimorphic; stipes with 1, 2, or 4 vascular bundles near base, adaxially grooved; lamina simple or 1-pinnate, with terminal pinnae similar to lateral pinnae; pinnae simple, entire, thickly papery to leathery, glabrous; veins reticulate, areoles without included free veinlets. Sori linear and forming a narrow longitudinal band between midrib and margin, or borne irregularly along veins, or converging to orbicular or oblong sori, or in long inframarginal grooves, or acrostichoid on abaxial surface of contracted fertile pinnae, exindusiate; paraphyses abundant, multicellular. Spores trilete, tetrahedral-globose, with tubercles or rodlets. n = 44, 108, 110, 114.
Sori broad, elongate, parallel to costa and edge, or more or less spreading along the veins.