Trees, unarmed. Raphides absent. Leaves opposite, usually with domatia; stipules caducous, interpetiolar or shortly united around stem, triangular to ovate. Inflorescences terminal or sometimes pseudoaxillary or appearing leaf-opposed due to anisophylly with 1 leaf caducous at subtending node, compound-cymose, several to many flowered, pedunculate or sessile and tripartite, bracteate. Flowers pedicellate, bisexual, monomorphic. Calyx limb campanulate, truncate to 5-dentate. Corolla yellow to white, salverform, inside villosulous in throat; lobes 5, convolute in bud, reflexed at anthesis. Stamens 5, inserted in corolla throat, exserted; filaments short; anthers dorsifixed. Ovary 2-celled, ovules 1 or usually 2-6 in each cell on axile placentas; stigma fusiform to cylindrical, exserted. Fruit black, baccate, subglobose, leathery or fleshy, with calyx limb deciduous; seeds 2-8, medium-sized, ellipsoid to subglobose.