Plants medium-sized to large, in thin to dense mats, light to dark green or yellowish, glossy. Stems creeping, , simple or sparingly and irregularly branched, ; hyalodermis absent, central strand sometimes present; pseudoparaphyllia foliose. Stem and branch leaves similar, erect to wide-spreading or squarrose, ovate, ovate-lanceolate, oblong-lanceolate, or oblong-ovate, , not or occasionally plicate; base not decurrent; margins plane or recurved, serrulate to entire proximally, serrate to serrulate distally; apex acute to subobtuse, sometimes acuminate; costa double and short or sometimes ecostate; alar cells differentiated, quadrate to rectangular; laminal cells smooth or prorulose at distal ends on abaxial surface; . Specialized asexual reproduction absent. Sexual condition dioicous [rarely synoicous], usually sterile; perichaetial leaves ± spreading, lanceolate to ovate-lanceolate, apex slenderly acuminate. Seta yellowish to red. Capsule cernuous, oblong-ovoid, straight to arcuate, somewhat contracted below mouth and wrinkled at neck when dry; annulus 2-seriate, persistent; operculum obliquely rostrate; peristome double; exostome teeth cross striolate proximally, papillose distally; endostome basal membrane high to low, segments keeled, cilia shorter than or same length as segments, in groups of 1-3. Calyptra naked. Spores globose to ovoid, smooth to minutely papillose.
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