Thesium goetzeanum Engl.

Species

Angiosperms > Santalales > Santalaceae > Thesium

Characteristics

Perennial herb with thick woody rootstock, sometimes with rhizomes; vegetative buds not very evident, ± 2 mm long, with triangular-acuminate scales 1–1.5 mm long; stems tufted from crown, 10–120 cm tall, sparingly to well branched, leafy; ribs strongly raised and extended into midribs of leaves and bracts.. Leaves mostly well-developed, with a few scales near base of stems, glaucous, thick-textured, ascending to somewhat spreading and then slightly recurved, linear-lanceolate, 6–20x1–1.  midway on stem, smaller upwards and passing into bracts, often shorter and slightly broader at base, apex apiculate, base broad, margins pale, cartilaginous, very minutely scabridulous or smooth, midrib raised on both surfaces.. Flowers 7–13 racemosely arranged at tips of branches, laxly panicled, usually solitary in each bract axil, occasionally paired or in 3-flowered cymules, sometimes all solitary except for 3-flowered terminal cymule; bracts usually adnate to entire length of pedicel, rarely only to lower half of peduncle, lanceolate, leaf-like, 4–12(–20)x1–2 mm; bracteoles 2, 2–7.5x0.4–1.3 mm; pedicels up to 0.5–2(–4) mm long; peduncles of cymules up to 10 mm.. Perianth greenish; tube 0.7–1.2 mm long, external glands ± conspicuous, elliptic, equalling tube; lobes triangular, 1.2–2x0.6–0.9 mm, subacute, densely white-bearded from apex.. Stamens inserted at base of perianth lobes; filaments 0.2–0.5 mm long, almost hidden by anthers; anthers 0.4–0.8 mm long.. Style 0.6–1.1 mm long; stigma reaching nearly to top of anthers.. Fruit yellowish buff or slightly reddish buff, ellipsoid-globose, 3–3.5 mm long, 2.5–3 mm in diameter, ribs strongly raised, reticulations rather weakly raised in comparison, becoming stronger only at final maturation.. Fig. 1/16 (page 4).
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Rhizomatous suffrutex, 0.1-0.5(0.8) m tall, vegetative scales present on rhizome and lower parts of aerial stems, stems 1 to 22(47), arising from rhizome at intervals, erect or suberect, virgate, often with vegetative shoots overtopping inflorescences, simple or occasionally branched, green, sulcate, moderately leafy (4 to 16 leaves per 50 mm at middle of stem). Leaves adpressed or slightly spreading, linear or narrowly obovate, 4.5-20.0 x 0.3-1.8 mm, apex acute or pungent and usually not cartilaginous, midrib raised on both surfaces, margins entire. Flowers usually solitary in bract axils, sometimes with simple 3-flowered dichasial cymes, or very rarely compound dichasial cymes, arranged in 4 to 13(19)-flowered monotelic racemose inflorescences, often terminating in simple dichasial cymes; cyme peduncles 0.5-6.0(10.5) mm long, pedicels 0-3(5) mm long. Bracts linear-lanceolate, 4.0-12.5 x 0.4-1.8 mm, usually acuminate, margins entire or occasionally scabrous, fused to entire pedicel or in cymes fused to about 1/3 of cyme peduncles; bracteoles 2.0-6.0 x 0.2-0.9 mm. Perianth 3.0-4.5 mm long, elongate receptacle absent, 'glands' often visible on outside; lobes narrowly triangular, 1.1-1.7 x 0.3-0.6 mm, apex hooded, with dense apical beard. Stamens inserted at base of tepals; filaments 0.3-0.4 mm long; anthers 0.4-0.7 mm long. Style 0.8-1.2 mm long, stigma ± opposite anthers. Placental column straight. Fruit sessile, 4.5-7.0 mm long including persistent perianth (2.5-4.5 mm long excluding perianth), 1.5-2.8 mm wide, with faint reticulate venation.
Stems few from the apex of a fairly stout woody rhizome, suberect, angular and sulcate, glabrous; branches suberect, sparingly leafy; leaves linear, acute or subacute, 3-7 lin. long, 1/2-3/4 lin. broad, thick and fleshy, with very finely scabrous cartilaginous margins, mostly with a distinct midrib or more or less keeled on the lower surface, glabrous; flowers few in leafy racemes, solitary in the bracts; bracts adnate to the peduncle, lanceolate or ovate-lanceolate, acutely acuminate, longer than the flowers, with distinctly cartilaginous scabrous margins; bracteoles similar to the bracts but about half as long; perianth urceolate, with distinct external glands, about 2 lin. long; segments linear-lanceolate, obtuse, 1-1/2 lin. long, hooded with a dense woolly apical beard; anthers included, reaching to the top of the perianth-tube, 3/8-1/2 lin. long; style 3/4 lin. long, reaching to nearly the top of the anthers; fruit ovoid-ellipsoid, 3 lin. long, conspicuously 10-ribbed, only very slightly reticulate between the ribs.
Stems several from a broad-topped woody rhizome, strongly ribbed and sulcate, glabrous; leaves broadly linear, acute, straight, 1/2-3/4 in. long, flat, rather thick and fleshy, becoming blackish when dry, glabrous; flowers solitary in the bract axils, subsessile, arranged in lax spikes; bracts as long as or longer than the open flower, linear-Ianceolate, acute, entire, glabrous; bracteoles about two-thirds as long as the bracts; perianth with large conspicuous external glands, about 1 1/2 lin. (3 mm) long; segments linear-lanceolate, subacute, 1 lin. (2 mm) long, infolded and deeply hooded, densely fringed and bearded to the base; anthers included towards the top of the perianth-tube, about 1/4 lin. long; style 1/2 lin. long, reaching to the top or the middle of the anthers; fruits ovoid-ellipsoid, 3 1/2 lin. long, strongly 10-ribbed, very slightly reticulate between the ribs.
Stems very short, up to 5 in. high, simple or forked, probably arising from a rhizome, slender, sulcate or angular, glabrous; leaves linear, with a cartilaginous acute apex, 4-5 lin. long, about 1/2 or 1/3 lin. broad, slightly keeled below, fairly thick and fleshy, glabrous; flowers few, solitary, axillary, very shortly pedunculate; bracts adnate to the apex of the peduncle, Iinear, with acutely subulate apex, equal to or slightly longer than the flowers, deeply concave on the upper surface, very minutely serrulate on the margin; bracteoles as long as the flower, narrower than the bracts; perianth white, 2 lin. long, without external glands; segments oblong-lanceolate, obtuse, 1 lin. long, hooded, flat, densely woolly-bearded; anthers partly included in the tube and partly exserted, 3/8 lin. long; style 1 1/8 lin. long, longer than the anthers.
Stems solitary or subsolitary from a small knotty rhizome, sparingly branched, erect, about 4 in. high, strongly ribbed, somewhat glaucous, glabrous; leaves few, linear, obtuse or subacute, 4-6 lin. long, 3/4-1 lin. broad, slightly keeled below, slightly glaucous, glabrous, thick and fleshy; flowers white, few in loose racemes, solitary in the bracts; bracts adnate to the short (up to 1 lin. long) peduncle, elliptic-lanceolate, obtuse, more or less boat-shaped, with smooth margins, longer than the flowers, entire; bracteoles about two thirds as long as the bracts; perianth 2 1/8 lin. long, with very large and conspicuous external glands; segments ovate-lanceolate, obtuse, 1 1/3 lin. long, with a dense woolly apical beard; anthers partly exserted, nearly 1/2 lin. long; style 1/2 lin. long, reaching to the base of the anthers; fruit not seen.
Suffrutex, caules numerosi, erecti, superne ramosi, 15-20 cm alti, conspicue angulati et sulcati. Folia inferne squamiformia, superne linearia vel lineari-lanceolata, acuta, 0.6-1 cm longa, apice brunnea. Infloresentia terminalis, racemosa; flores sessiles, singuli, rarius in cymas 3-floras in axillis bracteae dispositi; bracteae anguste ovato-lanceolatae, acutae, carinatae, carnosulae, 4 mm longae, floribus aequilongae, bracteolae 2, circiter 3 mm longae. Perianthium 3.5 mm longum, segmentis 2 mm longis elliptico-lanceolatis cucullatis apice dense barbatis. Antherae 0.75 mm longae. Stylus 1.25 mm longus, medio antherarum attingens. Fructus ovoideus, 3-4 mm longus, costis 10 conspicuis reticulationibus tenuibus instructus.
A subshrub; caudex small, thick; stems erect, 10-12 in. long, numerous, branched, glaucous, longitudinally grooved; leaves linear-lanceolate, apex cartilaginous, 7 lin. long, strongly keeled; bracts and bracteoles adnate to the pedicels; bracteoles 2-2 1/2 lin. long; perianth white, 1-2 lin. long; segments triangular, elongate, obtuse, 3/4-1 lin. long, margin inflexed, with a dense beard of long hairs at the apex and upper part of the margins; anthers 1/4 lin. long, equal in length to the filaments; style 3/8 lin. long; fruit ovoid, with thick prominent ribs, 1 3/4-2 lin. long, 1 1/4 lin. broad.
Few-stemmed, hemiparasitic, dwarf shrub, 0.1-0.3 m high; branches suberect, leafy. Leaves with midrib strongly keeled below, flat or cymbiform, linear in outline, up to 14 mm long, apex uncinate. Inflorescences of few-flowered, leafy racemes. Bracts leaf-like, ovate-lanceolate, adnate to peduncle. Flowers solitary in bract axils; green, white inside. Perianth urceolate, ± 4 mm long, with distinct external glands; segments linear-lanceolate, up to 3 mm long, hooded. Flowering time Oct.? Fruit ovoid-ellipsoid, ± 6 mm long; conspicuously 10-ribbed, slightly reticulate between ribs.
Dwarf shrub or shrub, 0.2-0.6 m high. Stems erect, many, branched, longitudinally grooved, glaucous. Leaves erect; blade linear-ovate, ± 14 mm long, strongly keeled, apex cartilaginous. Flowers: solitary in terminal raceme-like inflorescences; bracts adnate to pedicels, keeled, margins entire, glabrous; perianth bearded, 2-4 mm long, segments triangular-elongate, apices obtuse, 1.5-2.0 mm long, margins inflexed, dense beard of long hairs at apex and upper part of margins, white; Dec.-Apr. Fruit ovoid, 3-4 mm long, thick, ribbed.
Dwarf shrub, 0.12-0.30 m high. Stems simple or forked, slender, sulcate or angular; glabrous. Leaves fleshy; blade linear, 8-10 x 0.7-1.0 mm, apex acute and cartilaginous, glabrous. Flowers: few, solitary, axillary, in raceme-like inflorescences; bracts adnate to apex of peduncle, equal or slightly longer than flowers, rounded on back, minutely serrulate; perianth bearded, 4 mm long, segments narrowly ovate-oblong, apices obtuse, 2 mm long, hooded, densely woolly bearded, white; Dec.-Mar. Fruit ± 2-3 mm long.
Leaves ascending to somewhat spreading and then slightly recurved, 6–20(30) × 1–1.5(1.7) mm midway on stem, smaller upwards and passing into bracts, often shorter and slightly broader at base, linear-lanceolate, apex apiculate, base broad, margins pale, cartilaginous, very minutely scabridulous or smooth, thick-textured, midrib raised on both surfaces.
Flowers 7–13 racemosely arranged at tips of branches, laxly panicled, usually solitary in each bract axil, occasionally paired or in 3-flowered cymules, sometimes all solitary except for 3-flowered terminal cymule.
Perennial herb with thick woody caudex; stems tufted from crown, 110–300(600) mm tall, sparingly to well branched, strongly ribbed by raised vascular strands terminating in midribs of leaves and bracts, leafy.
Herb, up to 200 mm tall. Stems erect, overtopping branches few or many. Leaves 3-8 mm long, ascending. Flowers sessile or the lower on pedicels 1 mm long. Perianth lobes bearded.
Perianth greenish; tube 0.7–1.2 mm long, external glands conspicuous, elliptic, equalling tube; lobes 1.2–2 × 0.6–0.9 mm, triangular, subacute, densely white-bearded from apex.
Erect, much-branched herb, up to 200 mm tall. Leaves erect. Flowers in racemes. Bracts adnate to peduncles, margins entire, keeled. Perianth segments bearded. Flowers white.
Bracts 4–12(20) × 1–2 mm, lanceolate, leaf-like, usually adnate to entire length of pedicel, rarely only to lower half of peduncle; bracteoles 2, 2–7.5 × 0.4–1.3 mm.
Stout herb, up to 150 mm tall. Leaves straight. Perianth with segments flat, without hood. Bracts elliptic-lanceolate, obtuse. Flowers yellowish green.
Herb. Bracts scabrid at edges, longer than flowers. Perianth lobes bearded, longer than tube. Flowers solitary or in clusters of 2-4; white.
Fruit c. 3–3.5 × 2.5–3 mm, yellowish-or slightly reddish-buff, ribs strongly raised, reticulations rather weakly raised in comparison.
Herb, up to 200 mm tall. Perianth segments with deep hood, bearded. Bracts lanceolate, acute. Flowers greenish white.
Stamens inserted at base of lobes; filaments 0.2–0.5 mm long, almost hidden by anthers; anthers 0.4–0.8 mm long.
Pedicels (lowermost) 0.5–2(4) mm long; peduncles of cymules up to c. 10 mm.
Style 0.6–1.1 mm long; stigma reaching nearly to top of anthers.
Life form perennial
Growth form herb
Growth support parasite
Foliage retention evergreen
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.14 - 0.2
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light -
Soil humidity 1-3
Soil texture 7-8
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Distribution

Thesium goetzeanum world distribution map, present in Burundi, Botswana, Kenya, Lesotho, Mozambique, Malawi, Rwanda, eSwatini, Tanzania, United Republic of, South Africa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:780866-1
WFO ID wfo-0001134368
COL ID 7BWLD
BDTFX ID -
INPN ID -
Wikipedia (EN)
Wikipedia (FR)

Synonyms

Thesium coriarium Thesium deceptum Thesium goetzeanum Thesium macrogyne Thesium nigrum Thesium orientale Thesium rogersii Thesium caespitosum