Herbs, climbing. Roots tuberous. Stem and branches angular-sulcate, pubescent and villous. Petiole slender, 3-7 cm, pubescent and villous; leaf blade ovate-cordate, 6-11 × 4-9 cm, abaxially puberulent and setose on veins, adaxially densely setose, scabrous, basal lateral veins along margin of sinus, base cordate, margin irregularly serrate, apex shortly acuminate; sinus open, semicircular, 15-20 mm deep, 10-20 mm wide. Tendrils simple, pubescent. Male flowers 3-6 at apex of peduncle in sparse racemes; peduncle 2-5 cm, pubescent; pedicels filiform, 2-4 cm; calyx tube campanulate, ca. 4 × 4-5 mm; segments slightly reflexed, linear or linear-lanceolate, 4-12 mm, 3-veined, apex acuminate; corolla segments ovate-lanceolate, ca. 15 × 6 mm, apex acute; filaments filiform, 3-4 mm, puberulent; anthers oblong, ca. 2 mm. Female flowers solitary, larger than male flowers; pedicel slender, pubescent; calyx segments linear, 8-9 mm; corolla segments ovate-lanceolate, ca. 20 × 7 mm, distinctly 3-5-veined, apex shortly acuminate; ovary ovoid, ca. 7 × 6 mm, pubescent and villous, apex shortly rostrate; style ca. 5 mm, 3-fid; stigma enlarged, 2-lobed, reniform. Fruiting pedicel 2-3 cm, sparsely pubescent; fruit ovoid, 2-3 cm in diam., verrucose. Seeds broadly ovate, 6-7 × ca. 3.5 mm, ca. 1.5 mm thick, punctate, base slightly obtuse. Fl. and fr. Apr-Nov.