Plants green or yellow-green, appearing brown proximally, loosely foliose, some leaf bases exposed, only slightly comose at stem apex. Stems conspicuously tomentose. Leaves in loose, simple spirals or once circinate when dry, appearing firm or rigid, often fragile and erose, leaf tips often absent; inconspicuously undulate, especially when dry, fragile or not, erect-to erect-spreading when wet; undulate or plane; proximal laminal cells thin-walled and hyaline, sharply differentiated in shape and size from the papillose distal cells, which are papillose in the area of contact; leaves keeled in section, costa in apical region exposed adaxially by up to two stereid cells in width or completely covered with an adaxial epidermis of quadrate papillose cells; leaf cross section with 2-stratose areas beside the costa, the lamina irregularly 2-stratose in patches, lamina tattered, costa occasionally appearing undifferentiated in apical region of the leaf, adaxial stereid layer occasionally disappearing toward the apex, epidermal layer may be continuous throughout the leaf length.