Plants small, gregarious or in loose patches, dark green to blackish dry, green to light green wet; terricolous or saxicolous. Stems short, 2-5 mm high, sparingly branched; in section with central strand, inner cortical cells lax, outer cortical cells in 1-3 rows, smaller, reddish. Leaves incurled, spirally twisted or contorted dry, wide-spreading wet; shape extremely variable, oval to oblong on short-leaved plants or oblong to lingulate on long-leaved plants; apex acute to obtuse; margins narrowly revolute, entire. Costa percurrent, strongly convex ventrally in upper leaf; ventral superficial cells quadrate, papillose, dorsal superficial cells elongate, strongly incrassate, smooth; in section oval, guide cells 4-6, ventral cells in 1-2 layers, surface cells enlarged, frequently vertically elongated, papillose, dorsal stereid band 4-6 cells thick, dorsal surface cells substereids. Upper laminal cells quadrate to hexagonal, with 4-6 C-shaped papillae over lumen; cells of revolute margins smooth or nearly so, rarely all laminal cells only weakly papillose; basal cells rectangular, lax, smooth. Autoicous. Perichaetia terminal, leaves undifferentiated. Seta 6-15 mm long, yellow to reddish; capsule ovoid to cylindrical, 1.0-2.5 mm long, reddish brown; peristome teeth divided into 2 papillose filaments above short basal membrane, 0.3-0.5 mm high, yellowish, frequently irregular, broken; operculum conical to obliquely rostrate, 1 mm long; spores roundish, 17-25 µm, finely punctate.
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Leaves ovate to lingulate, apex broadly acute to rounded, apiculate or mucronate, margins revolute or occasionally plane, often bordered proximally with rectangular cells usually hidden in the revolute margins; costa percurrent or ending in apiculus or short mucro, with an adaxial pad of swollen cells similar to those of the lamina, distally broad, (4-)6(-8) cells across adaxial surface; distal laminal cells subquadrate, 9-13(-18) µm wide, µm, 1:1; strongly papillose. Sexual condition autoicous. Sporophytes exerted. Seta 0.6-1.2 cm. Capsule stegocarpic, not systylius, cylindric to ellipsoidal, erect and nearly straight, urn 0.9-1.5 mm; peristome commonly 210-390 µm, teeth of 32 filaments, weakly twisted, occasionally rudimentary, basal membrane 40-50 µm; operculum 0.5-0.7 mm. Spores 15-18 µm, broadly ellipsoidal, nearly smooth.
Plants gregarious or in loose patches. Stems 2-5 mm long, central strand present. Leaves incurled, spirally twisted or contorted when dry, oval to oblong or lingulate; apex acute to obtuse; margins narrowly revolute; costa percurrent, convex ventrally, ventral stereid band absent. Upper laminal cells quadrate to hexagonal, with 4-6 C-shaped papillae; basal cells lax, smooth. Capsule cylindrical, peristome teeth divided into filaments. Recognised by the enlarged ventral cells of the upper costa.
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