Leaves short-elliptic or long-obovate, sometimes ovate, apex very broadly acute to rounded, short-to long-awned, margins very broadly recurved or revolute to near apex, not bordered; costa usually long-excurrent, lacking an adaxial pad of cells, distally narrow, ca. 3-4 cells across the convex adaxial surface; distal laminal cells hexagonal, 10-15 µm wide, 1:1, densely papillose with 2-fid papillae. Sexual condition apparently dioicous but more probably rhizautoicous, occasionally autoicous. Sporophytes exerted. Seta length ca. 0.7-1.4(-2) cm. Capsule stegocarpic, not systylius, cylindric, erect and nearly straight, urn length ca. (1.7-)2.5-3.5 mm; peristome 800-1000 µm, teeth of 32 filaments twisted about one full turn, basal membrane 150-200 µm; operculum 1-1.5 mm. Spores ca. 10-13 µm, spheric, finely papillose.
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