Leaves elliptic, apex broadly acute to rounded-acute, mucronate or apiculate, margins recurved at mid leaf, bordered throughout or in lower 2/3 with 2(-3) rows of narrower, short-to long-rectangular, thicker-walled cells that are 1-2-stratose; costa percurrent or short-excurrent, lacking an adaxial pad of cells, distally narrow, 3-4 cells across the convex adaxial surface; distal laminal cells hexagonal to rectangular, (13-)16-24(-28) µm wide, 1-2:1, not or rarely very weakly papillose. Sexual condition autoicous. Sporophytes exerted. Seta 0.6-1.2 cm. Capsule stegocarpic, not systylius, short-cylindric and curved or ovoid and bilaterally symmetrical, nodding or occasionally nearly horizontal, urn 1-1.8 mm; peristome weak, ca. 100-200 µm, teeth 16, not twisted, divided nearly to the base into 2-3 branches and variously perforate, basal membrane very low or absent, or rudimentary or fragmentary and attached to the inside of the operculum; operculum 0.2-1.5 mm. Spores 25-35 µm, spheric, densely papillose.