Leaves ovate to oblong, apex broadly acute or occasionally rounded, apiculate or short-to long-mucronate, occasionally short-awned, margins recurved in middle 2/3 of leaf or occasionally almost plane, not or very weakly bordered distally with ca. 2 rows of rhomboid, less papillose cells; costa subpercurrent, percurrent or excurrent, lacking an adaxial pad of cells, distally narrow, 2-3 cells across the convex adaxial surface; distal laminal cells hexagonal, 15-20 µm wide, 1:1, densely papillose with several 2-fid papillae. Sexual condition autoicous. Sporophytes exerted. Seta 0.9-1.2(-1.9) cm. Capsule stegocarpic, rarely systylius, cylindric, erect and nearly straight, urn 1.5-2(-2.8) mm; peristome (150-)300-350 length µm, teeth 16, usually not twisted, divided to near the base or perforate basally, the teeth often basally fused in 2s or 4s, basal membrane very low; operculum 0.7-1 mm. Spores ca. 20-23(-25) µm, spheric or elliptic, densely papillose.
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