Leaves lingulate, apex rounded-acute, apiculate to short-mucro-nate, margins recurved from base to near the apex, not bordered; costa subpercurrent, percurrent or very short-excurrent, lacking an adaxial pad of cells but distally narrowing or weakly thickened, 3-4(-5) cells across the convex adaxial surface; distal laminal cells hexagonal, (12-)15-18 µm wide, 1:1, strongly papillose with 4-5 2-fid papillae. Sexual condition gonioautoicous. Sporophytes exerted. Seta 1.2-1.5(-2.5) cm. Capsule stegocarpic, not systylius, cylindric, erect and nearly straight, urn usually 3-4 mm; peristome length 1000-1500 µm, teeth of 32 filaments twisted at least one full turn, basal membrane 300-600 µm; operculum 1.3-1.6(-2) mm. Spores 11-15 µm, spheric, finely papillose.