Leaves obovate, elliptic or less commonly ovate, apex broadly acute to occasionally rounded, awned or long-mucronate, mar-gins strongly and broadly recurved in proximal 4/5 of leaf, not bordered; costa long-excurrent, with an elongate adaxial pad of bulging, rounded cells with 3-4 simple papillae, distally narrow, 2-4 cells across adaxial surface; distal laminal cells subquadrate, 12-16 µm wide, 1:1, smooth or less commonly papillose with 1(-2) scattered simple papillae. Sexual condition autoicous. Sporophytes exerted. Seta ca. 0.4(-0.9) cm. Capsule stegocarpic, not systylius, elliptic or cylindric, or erect and nearly straight, urn ca. 0.8-1.2 mm; peristome 100-150 µm, teeth 16, ligulate straight, or short, rudimentary, basal membrane lacking or very low; operculum 0.5 mm. Spores 15-18 µm, spheric, finely papillose.