Leaves ovate to ligulate, apex rounded or occasionally broadly acute, apiculate, margins strongly recurved or revolute in distal 4/5 of leaf, sometimes bordered distally with (2-)4 rows of thicker-walled cells, or occasionally border of rhomboidal, smooth cells commonly hidden in recurvature; costa ending in apiculus or occasionally percurrent or subpercurrent, lacking an adaxial pad of swollen cells or this poorly developed, distally not broadening, to 5-6 cells across the convex adaxial surface; distal laminal cells hexagonal or subquadrate, 10-13 µm wide, 1:1, strongly papillose with 2-fid or multiplex papillae. Sexual condition autoicous, but plants often bearing only antheridia or archegonia. Sporophytes exerted. Seta 0.5-0.8 cm. Capsule stegocarpic, cylindric or ovate, erect and nearly straight, urn 1-2.5 mm; peristome rudimentary or to 500 µm, teeth 16, irregular, divided to base and filamentous when long, straight or twisted, basal membrane very low or absent; operculum 0.7-1 mm. Spores 9-11(-15) µm, spheric, finely papillose or essentially smooth.
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