Trees; indumentum of simple or tufted hairs, sometimes with glandular hairs. Leaves opposite, entire, 3-5-plinerved; stipules 2. Inflorescences axillary, racemose. Plants dioecious. Inflorescence axillary, unbranched, male flowers 2 or 3 per bract. Male flowers: calyx closed in bud, valvate, 3-5-partite, reflexed; petals absent; disk absent; stamens 75-95; filaments free; anthers dorsifixed, 2-locular, connective sometimes widened; pistillode absent. Female inflorescence much shorter, 1-4-flowered. Female flowers: calyx 2-4-lobed, caducous; petals absent; disk absent; ovary 2-4-locular; ovules 1 per locule; styles 2-4, long, entire, conspicuously papillose. Fruit a drupe, indehiscent; endocarp crustaceous. Seeds ovoid; testa hard.