Female flowers: perianth as in the male but with smaller lobes and glands, ovary sessile, 1-locular with 3 parietal placentas borne high up on the ovary wall, each bearing near its base 1–3 ascending anatropous ovules.
Male flowers: perianth of 6–10 lobes in two rows, the inner slightly larger, disk glands opposite the outer perianth segments, stamens 9–12 in bundles of 3–4, alternating with the disk glands.
Flowers dioecious, with the males in axillary panicles and the females in spiciform racemes.
Leaves alternate, petiolate, 5–9-nerved from the base, margins serrate or dentate.
Stipules caducous, sometimes foliose.
Capsule 3-valved, 1–3-seeded.
Shrubs or trees.