veins and venules, the intervenal areas patulous reticulate; petioles to 1.5 cm long, 0.1-0.35 cm wide, tomentose; the stipules triangular, ca. 1 cm long, ca. 1.2 cm wide at base, acute, stiff, drying rubescent, with appressed white hairs. In-florescences axillary, to 9 cm long, the peduncles 1.0-3.5 cm long, 0.8-3.0 mm wide, simply terminating in a globose capitulum of flowers, or the axes with 2 pairs of branches, these opposite, perpendicular, well spaced, puberulent, each terminated by a rotund capitulum, ca. 2 cm in diam. Flowers yellowish; hypan-thium oblong, to 1.5 mm long, carnose, puberulent, the calycine cup urceolate, ca. 0.65 mm long, densely white villose within, pubescent outside, the 5 teeth disposed as points; corolla with the tube funneliform, ca. 5 mm long, ca. 0.6 mm wide, petaloid, puberulent inside and out, the lobes 5, reflexed, oblong rotund, 1.4-1.7 mm long, glabrous on the adaxial face; stamens 5, the anthers oblong, 1.1-1.5 mm long, sessile, attached near the mouth; ovary 2-loculate, the septum moderately thin, the style linear, far exserted, ca. 4.5-6.0 mm long, the stigmas clavate, 1.0-1.5 mm long. Fruits septicidally bivalvate, narrowly ovate oblong, to 6 mm long, the persistent calyx coroniform, ca. 1 mm long and wide, drying tan, vaguely longitudinally striate, covered with dense white hairs, the seeds linear fusiform, to 4 mm long, the body ca. 1 mm long, the wing narrower than the body, membranous, 1 end linear oblong, the other deeply cleft and biaristate.
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Vines, often in forest canopy, the stems quadrangular, minutely puberulent, armed with subnodal spines, the spines uncinate, stiff, lignose, to 2 cm long, 0.4-0.6 cm wide proximally. Leaves oblong or ovate oblong, 9-17 cm long, 4.3-9.0 cm wide, tapering acutely or occasionally widely deltoid or rounded at the apex, obtuse, truncate, often vaguely cordate at the base, the costa prominulous above, prominulous to prominent beneath, the lateral veins 8-10, arcuate, angular, char-taceous or papyraceous, usually discolorous, minutely pubescent on the veins above, beneath usually densely and finely golden puberulent with fewer larger hairs intermixed, or often sparsely puberulent, the hairs confined to the costa,