Plants small to medium-sized, in thin to dense mats, usually pale green to yellowish, sometimes dark green, often shiny. Stems creeping, freely branched, irregularly branched to pinnate; hyalodermis absent, central strand present or absent; pseudoparaphyllia linear to foliose. Leaves dimorphic; ventral usually smaller than dorsal, ± appressed, lanceolate to usually acuminate; dorsal leaves spreading to secund, oblong-ovate to ovate-lanceolate, somewhat asymmetric; base weakly decurrent; margins plane, entire or serrulate toward apex; apex acute or short-to long-acuminate, sometimes apiculate; costa double and short or ecostate; alar cells not or slightly differentiated; laminal cells smooth, . Specialized asexual reproduction absent. Sexual condition autoicous; perichaetial leaves erect, ovoid, oblong-ovoid, or oblong-ovate, apex subulate. Seta reddish, reddish orange, or yellow-orange. Capsule horizontal to pendent, ovoid to oblong-ovoid, ± asymmetric, constricted below mouth when dry; annulus differentiated; operculum apiculate to short-rostrate; peristome double; exostome teeth with external surface cross striolate, papillose distally; endostome basal membrane high, segments keeled, cilia usually in groups of 1-3. Calyptra naked. Spores spheric, finely papillose to nearly smooth.