Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, densely lanate when young, becoming sparsely so; tendrils bifurcate. Leaves simple; stipules brownish, oval or ovate-lanceolate, 3.5-8 × 2.5-4 mm, membranous, glabrous or subglabrate, entire, apex obtuse; petiole 0.5-4.5 cm, densely lanate, pubescent, or tomentose, becoming sparsely so; leaf blade ovate, 2.5-8 × 2-5 cm, basal veins 5, lateral veins 4-6 pairs, veinlets inconspicuous or slightly raised adaxially, base cordate or deeply so, notch rounded, margin with 9-16 obtuse teeth on each side or pinnatifidly divided. Panicle leaf-opposed, basal branches well developed or sometimes reduced to tendrils, rarely narrow and branches undeveloped; peduncle 0.5-2.5 cm, with arachnoid tomentum when young, becoming sparsely so. Pedicels 1.5-3 mm, glabrous. Buds obelliptic or globose, 1.5-2.2 mm, apex rounded. Calyx ca. 0.2 mm, glabrous, subentire. Filaments filiform, 1.5-1.8 mm; anthers yellow, oblong, 0.4-0.5 mm. Ovary elliptic-ovoid; style short, slender; stigma expanded. Berry purple-red at maturity, 5-8 mm in diam. Seeds obovoid, apex retuse, chalazal knot rounded or elliptic, ventral holes narrow, furrowed upward ca. 3/4 from base. Fl. Apr-Aug, fr. Jun-Oct.
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