Waltheria indica L.

Uhaloa (en)

Species

Angiosperms > Malvales > Malvaceae > Waltheria

Characteristics

Shrublet, subherbaceous at first, stems slender, hirsute, with stellate, tufted or simple bulbous based hairs, erect to spreading and bushy, 30 cm to 1 m tall, sometimes taller. Stipules linear-acute, 3-10 mm long. Leaves petioled; blade ovate, ovate-oblong, oblong to narrowly oblong, 2-10 cm long, 1-4.5 cm broad, crenate-dentate, rounded, subtruncate or broadly subacute at the apex, rounded, broadly cuneate or cordate at the base, nerves prominent beneath, laxly to densely stellate pubescent or hirsute with bulbous-based hairs on both surfaces; petiole hirsute, 0.6-3.5 cm long. Inflorescence axillary, cymose, usually tightly congested rarely (abnormal)? sublax, sometimes heads of flowers appear to be racemosely or paniculately disposed because the subtending leaves are much reduced or absent. Bracts usually 3 to a flower, linear acute, 3.5-5 mm long, differing slightly in width but mostly under 1 mm wide, dorsally hirsute. Calyx hirsute, 10-nerved; tube turbinate, about 2.5 mm long; lobes deltoid acute to deltoid acuminate, about 2 mm long. Petals yellow, ‘orange yellow’, turning reddish brown at maturity, oblong-cuneate to spathulate, about 4 mm long, 1.25 mm broad, dorsally sparsely hairy, adhering at the base to the stamen base, deciduous from the base but persisting for some time around the capsule, the lower portion held within the calyx-tube. Stamen opposite the petals, shorter than the full grown petals; filaments membrane-margined at the base or, more usually, almost to the apex, more or less united along the membrane edge, the apical unwinged filiform portion shortly and sparsely pubescent and at certain stages contorted; anthers erect or (especially in abnormal specimen) horizontal, cells parallel, subacute at both ends. Ovary 1-celled, obovoid slightly [as “slighly”] flattened, broadest at the apex, densely hirsute in upper portion; style excentric, about 3 mm long, sometimes rather contorted, fimbriate at the apex; ovules 2, ascending, anatropus. Capsule thin-walled, about 4 mm long, almost 2 mm broad at the apex, hirsute on upper portion, 1-seeded or rarely the second ovule developing as well; seed obovoid; embryo straight.
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Herb or small shrub 0.60-2 m. high, the young stems grayish and soft-velvety-lanate-arachnoid. Leaves with a pedicel 0.5-3.3 cm. long, lanate-arachnoid to sericeous, the stipules subfiliform, the blade narrowly ovate to ovate, rounded or truncate or somewhat subcordate at the base, obtuse or subacute at the apex, 3.5-12 cm. long and 1.5-7 cm. wide, chartaceous, the margins serrate-crenate, grayish and soft-velvety-lanate-arachnoid on both sides, the indumentum slightly denser on the lowger surface, the nervation not reticulate, impressed above and prominent below. Inflorescences axillary or terminal, of tight-globose, subsessile to long-pedunculate glomerules, the peduncles lanate-arachnoid to sericeous. Flowers sessile, the bracts linear-lanceolate, about as long as the flowers, sericeous; calyx tubular-campanulate, somewhat 5-angled, ca. 4-4.5 mm. long, divided about 112 way into 5 triangular-subulate lobes ca. 2-2.5 mm. long, long-sericeous outside, glabrous inside except the sericeous lobes; petals obovate-oblong, unguiculate, 4-4.5 mm. long and 1-1.5 mm. wide, yellow or orange, glabrous outside, scatteringly arachnoid near the base just above the claw inside; staminal tube ca. 2 mm. long, glabrous; anthers subsessile, ca. 0.8 mm. long; ovary oblong-ellipsoid, ca. 1.5 mm. long and 0.75 mm. wide, sericeous on the upper half; style geniculate in the middle, ca. 1.5 mm. long, loosely arachnoid; stigma plumose-papillate. Capsule included in the calyx and surrounded by the marcescent corolla, obovoid, ca. 2.5-3 mm. long and 1.5-2 mm. wide, sericeous above, 1-seeded; seed obovoid, ca. 1.8-2.1 mm. long and 1-1.3 mm. wide, the testa smocth.
Plants branched or nearly simple, to 7–12 dm, indument variable, tomentose to short-woolly, sparsely to densely pubescent. Stems to 8 mm wide, nodes even or extended to 0.5 mm. Leaves: stipules linear-triangular, 3–6 mm; petiole 2.5–5 cm × 1–4 mm; blade concolor or discolor, drying olive, ovate-oblong to lanceolate, to 7(–10) × 5 cm, base subcordate to obtuse, margins crenate-serrate, serrate, irregularly serrate, or crenate-dentate, apex narrowly rounded, obtuse, or subacute, surfaces tomentose or pubescent, trichome rays 0.4–1 mm, and sparsely hispidulous, slightly scabrous or not. Inflorescences subsessile, or on short pedunclelike lateral branches to 3.6 cm, glomerules 5–20 along stems, compact, often dense, sympodial, at times lax clusters with terminal flower and subtending dichasia; bracteoles free or with 1 adherent to stipule forming an elliptic 2-cuspidate foliole at glomerule base, bracts subequal, lanceolate or linear, 0.3–1.5 mm wide, apex acute, entire. Flowers sessile or subsessile; calyx 4.2–setose apically; styles 0.9–1.7 × 0.1–0.3 mm, lateral or excentric, broader and stellate-hirsute distal to base; stigmas 12-branched, plumose (1.2–1.4 × 0.5–0.8 mm, column 0.9–1.2 mm, branches to 0.7 mm) or dissolute-penicillate (0.6–1.2 × 0.7–1.6 mm, column 0.4–0.7 mm, branches to 1.1 mm). Capsules oblique, obconic, or obovoid, 2.1–3 × 1.5–2 mm, walls with endocarp corneous for 1+ mm from apex, sometimes merely membranous along one or both valve margins laterally; partially apically dehiscent to 2-valvate. Seeds dark brown, obovoid or obconic, 2–2.2 × 1.2–1.5 × 1–1.3 mm, smooth. 2n = 24, 26 (India), 40 (Africa).
Erect or ascending subshrubs, up to 1 m tall, multi-branched. Branchlets densely puberulent. Petiole 0.5-1 cm; leaf blade ovate or long elliptic-ovate, 2.5-4.5 × 1.5-3 cm, both surfaces densely puberulent, base rounded or shallowly cordate, margin serrulate, apex obtuse. Inflorescence cymose, capitate, axillary; peduncle nearly absent to 1.5 cm. Epicalyx lobes nar-rowly lanceolate, ca. 4 mm. Calyx tube-shaped, 5-lobed, 3-4 mm, lobes triangular, much longer than tube. Petals yellowish, spatulate, apex truncate, slightly longer than sepals. Filament tube enclosing ovary. Ovary sessile, puberulent; style inserted obliquely; stigma fimbriate. Capsule obovoid, ca. 3 mm, hairy, 1-seeded, enclosed by persistent calyx. Seed obovate, very small. Fl. summer-autumn.
A herb or shrub. It grows 4 m high. The leaves are simple and 2-10 cm long by 1-4 cm wide. There are white stellate hairs on both sides. The flowers are in groups in the axils of leaves or at the ends of branches. The flowers are 3-4 mm wide. They have 5 yellow or orange petals. The fruit is a capsule. They are brown and 2 mm long and have 1 valve. They are tightly clustered together and are hairy. There is 1 seed.
Flowers yellow, in dense subsessile or long-pedunculate, axillary, globose heads, or the upper inflorescences forming a short interrupted or continuous spike, or irregularly collected into dense, leafy or leafless cymes or corymbs; bracts and bracteoles about as long as the flowers, linear or linear-lanceolate, tomentose outside.
Leaf-lamina up to 9 × 4 cm., ovate-oblong to oblong, apex obtuse or subacute, margin serrate-crenate, base rounded and 5-nerved, nervation somewhat impressed above, prominent below; petiole up to 3·3 cm. long.
Stamens with the filaments united into a tube shorter than the petals; anther-thecae parallel, bluntly mucronate at base and apex.
Petals as long as or slightly longer than the calyx, obovate-oblong, shortly clawed, glabrous or the apex stellate-ciliolate.
Shrublet, up to 1 m high. Flowers congested in axillary glomerules. Petals 4.00 x 1.25 mm. Calyces hirsute. Flowers yellow.
Ovary oblong-ovoid, hirsute in the upper half; style pubescent, somewhat excentric, c. 1·5 mm. long; stigma penicillate.
Bushy herb usually 0·6–1·3 m. tall, rather sparsely stellate-pubescent to stellate-tomentose in its vegetative parts.
Calyx 2–3 mm. long, campanulate, villous, divided somewhat less than 1/2 way into 5 triangular lobes.
Capsule c. 3 × 2 mm., obovoid, villous above, 1-or rarely 2-seeded.
Seeds c. 2 × 1·4 mm., dark brown, obovoid; testa smooth.
Petals yellow, turning orange or brown with age
Erect herb or shrub to 7 ft. high
Life form perennial
Growth form
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 1.0 - 1.1
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

A tropical plant. It grows on sandy shores. It grows in monsoon and vine forests throughout the tropics. It also grows in woodland and savannah. In tropical Queensland it grows from sea level to 800 m altitude. In Argentina it grows from sea level to 1,000 m above sea level. In Yunnan.
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Woodland and bushland; often on rocky hills or on gravelly soil; river-beds; humid sand of depressions; dune and sandy, grassy plains; flooded areas; also ruderal; savannah; cultivated ground; at elevations up to 1,650 metres in Africa.
Common in open places.
Light -
Soil humidity 1-3
Soil texture 7-8
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses animal food fiber food material medicinal social use vertebrate poison
Edible fruits
Therapeutic use Dietary Aid (bark), Respiratory Aid (bark), Strengthener (bark), Throat Aid (bark), Dietary Aid (leaf), Respiratory Aid (leaf), Strengthener (leaf), Dietary Aid (root), Respiratory Aid (root), Strengthener (root), Throat Aid (root), Syphilis (root), Laxative (unspecified), Pediatric Aid (unspecified), Pulmonary Aid (unspecified), Respiratory Aid (unspecified), Abortifacient (unspecified), Collyrium (unspecified), Cough (unspecified), Emollient (unspecified), Eruption (unspecified), Eye (unspecified), Fever (unspecified), Giddiness (unspecified), Hemoptysis (unspecified), Preventitive(Fever) (unspecified), Skin (unspecified), Soap (unspecified), Sore (unspecified), Sterility (unspecified), Sudorific (unspecified), Vertigo (unspecified), Enterorrhagia (unspecified), Purgative (unspecified), Syphilis (unspecified), Dentifrice (unspecified), Diarrhea (unspecified), Wound (unspecified), Asthma (unspecified), Debility (unspecified), Restorative (unspecified), Depurative (unspecified), Tea (unspecified), Sore(Throat) (unspecified), Weight-Loss (unspecified), Abortifacient agents (unspecified), Antipyretics (unspecified), Cathartics (unspecified), Emollients (unspecified), Abortifacient agents (whole plant), Antipyretics (whole plant), Cathartics (whole plant), Cough (whole plant), Emollients (whole plant), Hemorrhage (whole plant), Syphilis (whole plant)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Waltheria indica habit picture by Maarten Vanhove (cc-by-sa)
Waltheria indica habit picture by Nelson Zamora Villalobos (cc-by-nc)
Waltheria indica habit picture by Trap Hers (cc-by-sa)

Leaf

Waltheria indica leaf picture by Leonardo Hirata (cc-by-sa)
Waltheria indica leaf picture by julian spezz (cc-by-sa)
Waltheria indica leaf picture by Mathias Antoniol (cc-by-sa)

Flower

Waltheria indica flower picture by Tna (cc-by-sa)
Waltheria indica flower picture by Tna (cc-by-sa)
Waltheria indica flower picture by Mathias Antoniol (cc-by-sa)

Fruit

Waltheria indica fruit picture by Nelson Zamora Villalobos (cc-by-nc)

Distribution

Waltheria indica world distribution map, present in Angola, Anguilla, Argentina, Antigua and Barbuda, Australia, Burundi, Benin, Bonaire, Sint Eustatius and Saba, Burkina Faso, Bangladesh, Bahamas, Belize, Bolivia (Plurinational State of), Brazil, Barbados, Botswana, Central African Republic, China, Côte d'Ivoire, Cameroon, Congo, Colombia, Comoros, Cabo Verde, Costa Rica, Cuba, Cayman Islands, Djibouti, Dominica, Ecuador, Egypt, Eritrea, Ethiopia, Fiji, Ghana, Guinea, Guadeloupe, Gambia, Guinea-Bissau, Equatorial Guinea, Grenada, French Guiana, Guam, Guyana, Honduras, Jamaica, Kenya, Cambodia, Lao People's Democratic Republic, Liberia, Saint Lucia, Sri Lanka, Madagascar, Mexico, Myanmar, Mozambique, Mauritania, Montserrat, Martinique, Mauritius, Malawi, Namibia, Niger, Nigeria, Nicaragua, Panama, Peru, Philippines, Puerto Rico, Paraguay, Réunion, Rwanda, Saudi Arabia, Sudan, Senegal, Singapore, Sierra Leone, Somalia, South Sudan, Suriname, eSwatini, Seychelles, Turks and Caicos Islands, Chad, Togo, Thailand, Tonga, Trinidad and Tobago, Taiwan, Province of China, Tanzania, United Republic of, United States Minor Outlying Islands, United States of America, Saint Vincent and the Grenadines, Venezuela (Bolivarian Republic of), Viet Nam, Yemen, South Africa, Zambia, and Zimbabwe

Conservation status

Waltheria indica threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:320781-2
WFO ID wfo-0000427365
COL ID 7G5QT
BDTFX ID 166582
INPN ID 447631
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Waltheria laxa Waltheria americana Waltheria erioclada Waltheria prostrata Waltheria africana Waltheria angustifolia Waltheria arborescens Waltheria corchorifolia Waltheria elliptica Waltheria guineensis Waltheria wildii Waltheria makinoi Waltheria martii Waltheria americana var. elliptica Waltheria americana var. glandulosa Waltheria americana var. densiflora Waltheria americana var. indica Waltheria americana var. subspicata Waltheria indica var. americana Waltheria indica var. prostrata Waltheria indica