Plants green, yellow-green, brownish, brown-red, or rarely clear red in extremely exposed habitats. Stems with axillary hair distal cells 1-4. Stem leaves narrowly ovate to triangular-ovate, falcate or sometimes straight, concave or slightly so; margins denticulate, sometimes only in part or indistinctly; apex acuminate, not incurved; costa to 1/2-4/5 leaf length, occasionally double or shorter; alar region narrowly transversely triangular, supra-alar cells small, not forming region with alar cells.