Plants small, yellow-green, dark green, or brownish, glossy. Stems 1-2 cm, not complanate-foliate, irregularly to regularly pinnate to 2-pinnate. Leaves closely imbricate to slightly spreading, ovate, ; margins serrulate; apex gradually acuminate, often abruptly piliferous; ecostate or costa double; alar cells elongate, inflated, pigmented, walls thin, region of 1-4 cells, middle lamella not apparent, supra-alar cells differentiated, ; laminal cells vermicular, smooth or 1-seriate multipapillose. Sexual condition dioicous or phyllodioicous. [Seta 1-2 cm. Capsule inclined to horizontal, ovoid to cylindric; exothecial cell walls not or slightly collenchymatous; operculum short-to long-rostrate or conic].
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