Perennial herbs or subshrubs, with taproot; stems erect, arcuate or sometimes prostrate and rooting at nodes. Leaves simple, opposite, petiolate, ovate or deltate to linear, sometimes trilobed to pinnately lobed (rarely ternately compound), entire or remotely to closely serrulate or serrate, 3-nerved; base attenuate or truncate. Capitula terminal on main or lateral shoots, solitary or few, long-pedunculate, radiate; involucral bracts in c. 2 subequal series, all lanceolate to ovate, herbaceous; receptacle paleate; paleae (ob-)lanceolate to ovate or oblong, rigid, keeled, obtuse or subapiculate, ± hooded, dorsally scabrous. Ray florets female, fertile; ligule 2-or 3-lobed, yellow. Disc florets bisexual, fertile, rarely male; corolla 5-lobed, yellow; anther sacs black. Achenes usually obovoid to obconical, truncate, sometimes shortly and irregularly winged, apically shortly pilose; ray achenes 3-angled; disc achenes 2-or 4-angled. Pappus of 1–6 short fragile awns, a ring of tiny scales, or absent.
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