Plants to 2 cm. Stem leaves irregularly twisted or curved, loosely spiraled around stem and ± undulate when dry, spreading to wide-spreading, sometimes subsquarrose, recurved when moist, oblong-lanceolate to broadly lanceolate, 0.7-2 mm; margins entire to ± sharply serrulate near apex; apex sharply acute, apiculus usually present; costa ending near apex, abaxial surface smooth or covered by cells only at apex; basal laminal cells short-rectangular; distal cells 5-11 µm, papillae 3-6 per cell, walls somewhat thick. Specialized asexual reproduction by clavate to elliptic gemmae, longitudinal walls usually absent. Sexual condition synoicous or autoicous. Spores 17-22 µm.