Plants medium-sized, caespitose, yellowish green or olivaceous above, brown to dark brown below; saxicolous to terricolous. Stems to 40 mm tall, branching dichotomously or by subperichaetial or subperigonial innovations, sparsely tomentose below; rhizoids smooth to coarsely papillose, red-brown or red; in section pentagonal to subround, inner cortex 7-14 cells across, thin-walled to incrassate, smaller towards outside, outer cortical cells smaller, in 1-4 rows, substereids or stereids. Leaves ± crowded, ± equal in size, erect to appressed when dry, widespreading to squarrose-recurved from ± sheathing bases when wet, in 5 indistinct rows, spiralled around stem; ovate-lanceolate to lanceolate or oblong-lanceolate, 1.4-3.0(-3.6) mm long; ventral surface keeled; apex acute to acuminate, toothed; base ovate to oblong, ± sheathing, margins decurrent; margins plane, serrate to strongly serrate in upper half to two-thirds, teeth frequently multicellular, unistratose with bistratose patches, thickened. Costa ending below apex to subpercurrent, ventral and dorsal superficial cells rounded to elongate, incrassate; in section round, bulging dorsally, lamina ventrally inserted, not differentiated or 2-4 ventral guide cells larger, ventral stereid band absent, dorsal cells in 1-3 rows, incrassate to stereids, with dorsal teeth distally, teeth multicellular. Upper laminal cells small, irregularly rounded-hexagonal, flat, (7.0-)8.5-12.5(-14.0) µm, smooth to papillose, 1-4 papillae scattered over lumen, small, low; basal cells rhomboidal to rectangular, thin-walled to incrassate, smooth or walls papillose. Gemmae axillary, sub-clavate, uniseriate, with transverse septa. Dioicous. Perigonia terminal, leaves broadly ovate to orbicular. Perichaetia terminal; inner leaves oblong-acuminate or ovate-acuminate to lanceolate, 1.2-2.5(-2.8) mm long; apex acuminate; margins plane, entire to crenulate above; costa frequently weak, ending below apex; laminal cells rounded to vermiculate above, vermicu late below, occasionally papillose. Seta 3-5 mm long, yellowish or brownish. Capsule exserted, erect, elliptical or narrowly pyriform, 8-ribbed, urn 1.0-1.8 mm long, neck 0.3-0.6 mm long; exothecial cells rectangular to quadrate, irregularly incrassate, smaller at mouth, rounded-quadrate, cells of ribs differentiated; annulus deciduous. Peristome double, recurved when dry; exostome teeth in 8 pairs, narrowly triangular or lanceolate, fused below, perforated above, 180-300 µm long, densely papillose above, striolate-papillose below, yellowish or yellowish brown; endostome segments alternating with teeth pairs, linear, minutely papillose or smooth, pale yellow or hyaline, basal membrane low or absent. Operculum 0.5 mm long, oblique. Calyptra 2.5-3.0 mm long, smooth, naked. Spores 15.0-22.5 µm.