Plants to 2.3 cm. Stem leaves loosely erect and ± spirally twisted to somewhat contorted, sometimes falcate-secund at stem apices when dry, erect-spreading to spreading when moist, lanceolate, narrowly so, elongate-lanceolate, oblong-lanceolate, or linear-lanceolate, 1-2 mm; margins entire or irregularly serrulate or serrate near apex, somewhat wavy; apex acute with gradual apiculus, apiculate, short-apiculate, or short-acuminate; costa ending below apex, abaxial surface covered by quadrate papillose cells distally; basal laminal cells subquadrate to rectangular; distal cells 7-11 µm, papillae 2-4 per cell, walls somewhat thick. Specialized asexual reproduction by clavate, elliptic, or subcylindric gemmae, longitudinal walls present or absent. Sexual condition dioicous. Sporophytes unknown.