Leaf blades: margins usually dentate to laciniately pinnatifid, rarely entire, faces mostly glabrous and often glaucous. Peduncles mostly longer than leaves at flowering, glabrate or apically ± villous to lanate. Phyllaries ± stramineous proximally, green, often purple-black blotched or spotted, or with a purple-black midstripes, rarely nearly all black, usually ovate or obovate, sometimes lanceolate, subequal to unequal at flowering, margins ± ciliate, especially distally, faces glabrous or slightly pubescent basally or medially; outer mostly glabrous adaxially. Corollas usually orange or yellow, sometimes pinkish, subequal to or surpassing inner phyllaries. Cypselae: bodies ± gradually tapered to beaks, ribs ± weakly and uniformly ridged (not thickened distally). 2n = 18, 34, 36.