Trees; dioecious; twigs and foliage glabrous or minutely pubescent with sim-ple hairs. Leaves alternate, petiolate, exstipulate; blade triplinerved, biglandular at base beneath, the margins entire to remotely crenulate. Inflorescences axillary (more or less fascicled), spiciform; c flowers in glomerules, 9 flowers solitary at each node; bracts inconspicuous, eglandular. Staminate flowers subsessile or pedicellate; calyx splitting at anthesis into 3 or 4 valvate lobes; petals absent; disc massive, hirsu-tulous; stamens 5-8 (mostly 6), the filaments free, the anthers dehiscing in-trorsely and longitudinally, the outer locule-halves larger than the inner, with enlarged glandular connective; pollen grains subglobose (slightly oblate), 3-col-porate, tectate (psilate); pistillode 4 3-lobed. Pistillate flowers pedicellate; calyx-lobes 4 or 5, slightly imbricate; petals absent; disc annular, pulviniform, hirsu-tulous (? confounded with the base of the ovary); ovary of 3 carpels, the ovules 1 per locule, the styles free, short, recurving, undivided, papillate. Fruits capsular, the columella persistent, massive; seeds flattened, ecarunculate, the outer coat fleshy, the inner coat reticulate-striate, the cotyledons broad and thin.
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