Alstonia scholaris (L.) R.Br.

Indian devil tree (en)

Species

Angiosperms > Gentianales > Apocynaceae > Alstonia

Characteristics

Medium-sized to big tree 10-50(-60) m high, 20-80(-130) cm dbh, fluted at the base or forming tall and steep buttresses up to 10 m high, spreading for up to 4 m at the base. Bark smooth, scaly or shallowly fissured and peeling off in rectangular flakes, fawn or light brown; inner bark granular, creamy, yellow or straw-coloured, with copious white latex. Branchlets glabrous. Leaves in whorls of 4-8(-9); petiole 5-20(-25) mm long, slightly winged, with intrapetiolar stipule at the base and 1-3 mm long, colleters narrowly triangular and densely packed together in the axils; blade narrowly elliptic to obovate, (5-)6-17(-22) by (1.5-)2.5-7.5(-8.5) cm, 1.7-4.5 times as long as wide, apex obtuse or rounded or often retuse (in young trees, the leaves usually much larger, shortly acuminate and with blunt acumen up to 1 cm long), base often decurrent onto the petiole, less often acute or obtuse, glabrous or velutinous beneath; 25-45(-55) pairs of secondary veins, rather straight, forming an angle of (70-)80-90° with the midrib, (1.5-)2-5(-7) mm from each other; tertiary venation reticulate, sometimes admedial ramified, conspicuous above. Inflorescence 4-13(-17) cm long, many-flowered, mostly formed of two dense bunches of flowers; pedicels 0-2 mm long, pubescent. Bracts and bracteoles scale-or sepal-like, ovate or narrowly ovate, acuminate or sometimes trilobed or irregularly lobed at the apex, 1-3 mm long, ciliate. Flowers fragrant. Sepals pale green, connate at the base for 0.3-0.7(-1) mm, ovate, 1.5-2.4 by 0.8-1.9 mm, obtuse, sometimes acute or mucronulate, pubescent outside, minutely pubescent or less often glabrous inside, erect, ciliate. Corolla lobes sinistrorse; white, yellow or cream, 7-12 mm long in the mature bud and forming an ovoid head, 2.5-3.5 by 1.7-2.4 mm; pubescent or partly pubescent outside (frequently glabrous on basal half, at least within the calyx); tube almost cylindrical, slightly widening around the anthers and at the base around the ovary, 7-10 by 0.8-1.6 mm; lobes broadly ovate or suborbicular, 3-5 by 2.5-4.5(-7) mm, 0.6-1.1 times as long as wide, pilose inside. Stamens inserted at 5.2-8.4 mm from the base; anthers ovate, 0.9-1.2 by 0.4-0.5 mm, obtuse. Pistil 6-9.5 mm long; ovary ovoid or broadly ovoid, 0.9-1.5 by 0.7-1 mm, densely pubescent, with or without a narrow disk-like thickening at the base; style 4.2-8 mm; style head pagoda shape, 0.5-1 mm high, glabrous, with an elongate and robust cleft stigmoid apical part 0.3-0.7 mm high. Fruit a pair of follicles, 20-40(-63) cm by c. 2(-2.5) mm, glabrous. Seeds dark brown, oblong, 4.5-5.5(-7) by 1.2-1.6(-2) mm, glabrous on both sides, ends rounded; margin slightly thickened, longest cilia 8-12(-14) mm long, becoming much shorter or glabrous towards the margins.
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Tree to 33 m high. Leaves 4–7-whorled; petiole 7–12 mm long; lamina elliptic to elliptic-lanceolate or elliptic-obovate, 11.5–23 cm long, 4–7.5 cm wide, cuneate, shortly acuminate, rounded or dentate at apex, discolorous; secondary veins 30–40 each side of midrib, 2–6 mm apart; tertiary venation obscure. Inflorescence to 12 cm long. Flowers 7–10 mm long, 2.8–10 mm diam.; pedicels 0.8–1.3 mm long. Sepals lanceolate-ovate, 1.8–2.2 mm long. Corolla cream to white; tube 5–9 mm long, with sparse to dense indumentum at top; lobes lanceolate, 1.5–4.3 mm long, with sparse to dense indumentum internally and externally. Fruit 15–32 cm long. Seeds oblong, 4–5 mm long; hairs 7–13 mm long.
Trees to 40 m tall, glabrous. Bark gray; branchlets copiously lenticellate. Leaves in whorls of 3-10; petiole 1-3 cm; leaf blade narrowly obovate to very narrowly spatulate, 7-28 X 2-11 cm, leathery, base cuneate, apex usually rounded; lateral veins 25-50 pairs, at 80-90° to midvein. Cymes dense, pubescent; peduncle 4-7 cm. Pedicel usually as long as or shorter than calyx. Corolla white, tube 6-10 mm; lobes broadly ovate or broadly obovate, 2-4.5 mm, overlapping to left. Ovaries distinct, pubescent. Follicles distinct, linear, to 57 cm X 2-5 mm. Seeds oblong, margin ciliate, ends with tufts of hairs 1.5-2 cm. Fl. Jun-Nov, fr. Oct-Dec. 2n = 22, 44*.
A tall tree. It grows 40 m tall. It has buttresses. The bark is brown and cracked. The leaves are spaced along the branches and are in rings. The leaves are entire and oval. They are dark green and blue-green underneath. The flowers are in the axils of leaves near the ends of branches. The flowers have a sweet scent. The fruit follicles are 30 cm long. The fruit are yellow to green and not fleshy. The seeds are about 5 mm long. They are rounded at both ends.
Life form perennial
Growth form tree
Growth support free-standing
Foliage retention evergreen
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 37.5 - 40.0
Root system -
Rooting depth (meter) -
Root diameter (meter) 0.5
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

It is a subtropical plant. It grows in the wet zone. It grows in coastal areas in Papua New Guinea. In southern China it grows between 200-1,000 m above sea level. In XTBG Yunnan.
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A dominant canopy species found in coastal mesophyll vine forest with a canopy height of 35-42 metres, in palm-dominated forests and in notophyll vine forests.
Grows in notophyll vine forestsin a variety of soils derived from volcanic substrates.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 10-11

Usage

UsesThis species is the most important source of the pulai timber. This timber is used for household items and the pulp makes a high quality paper. As it is so light it can be used for fishing net floats. It can be used for the treatment of diarrhoea and stomach ache, snake bites, and for treating Herpes zoster by applying the latex on the affected parts. The tree is also sometimes used as an ornamental.
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The bark is used as a spice and to flavour liqueurs. The latex is used for chewing gum.
Uses animal food charcoal construction dye environmental use essential oil fiber food fuel gum invertebrate food material medicinal ornamental poison social use spice timber wood
Edible barks fruits leaves saps
Therapeutic use Abdominal neoplasms (bark), Abdominal pain (bark), Abscess (bark), Anthelmintics (bark), Anti-bacterial agents (bark), Antidepressive agents (bark), Anti-infective agents (bark), Anti-inflammatory agents (bark), Antineoplastic agents (bark), Antipruritics (bark), Antipyretics (bark), Antirheumatic agents (bark), Appetite stimulants (bark), Arthralgia (bark), Asthenia (bark), Asthma (bark), Astringents (bark), Bronchitis (bark), Cardiotonic agents (bark), Cardiovascular system (bark), Colic (bark), Diarrhea (bark), Dysentery (bark), Dyspepsia (bark), Ear diseases (bark), Fever (bark), Filariasis (bark), Galactogogues (bark), Heart diseases (bark), Helminthiasis (bark), Hypophosphatemia, familial (bark), Hypotension (bark), Lactation disorders (bark), Leishmaniasis (bark), Leprosy (bark), Leukorrhea (bark), Malaria (bark), Neoplasms (bark), Pain (bark), Parasympatholytics (bark), Pneumonia (bark), Postnatal care (bark), General tonic for rejuvenation (bark), Skin diseases (bark), Snake bites (bark), Tuberculosis (bark), Ulcer (bark), Urticaria (bark), Vitiligo (bark), Wounds and injuries (bark), Antiperiodic (bark), Depurative (bark), Antidepressive agents (flower), Anthelmintics (fruit), Epilepsy (fruit), Mental disorders (fruit), General tonic for rejuvenation (fruit), Syphilis (fruit), Antiperiodic (fruit), Anti-infective agents (leaf), Antiviral agents (leaf), Beriberi (leaf), Diarrhea (leaf), Dysentery (leaf), Edema (leaf), Fever (leaf), Hypotension (leaf), Malaria (leaf), Skin diseases (leaf), Snake bites (leaf), Tuberculosis (leaf), Ulcer (leaf), Urticaria (leaf), Wounds and injuries (leaf), Abdominal pain (plant exudate), Antineoplastic agents (plant exudate), Antirheumatic agents (plant exudate), Cough (plant exudate), Earache (plant exudate), Disorder of ejaculation (plant exudate), Gonorrhea (plant exudate), Neoplasms (plant exudate), Toothache (plant exudate), Ulcer (plant exudate), Wounds and injuries (plant exudate), Anthelmintics (rhizome), Antineoplastic agents (root), Appetite stimulants (root), Leukorrhea (root), Liver diseases (root), Stomach diseases (root), Earache (stem), Leukorrhea (stem), Headache (unspecified), Ache(Leg) (unspecified), Antidote(Antiaris) (unspecified), Asthma (unspecified), Astringent (unspecified), Beriberi (unspecified), Boil (unspecified), Cancer (unspecified), Cholera (unspecified), Cough (unspecified), Diabetes (unspecified), Diarrhea (unspecified), Dyschezia (unspecified), Dysentery (unspecified), Enterosis (unspecified), Expectorant (unspecified), Fever (unspecified), Hemiplegia (unspecified), Internal (unspecified), Malaria (unspecified), Nausea (unspecified), Neuralgia (unspecified), Phthisis (unspecified), Piles (unspecified), Pneumonia (unspecified), Poison(Arrow) (unspecified), Rheumatism (unspecified), Stomachic (unspecified), Tonic (unspecified), Toothache (unspecified), Tumor (unspecified), Vermifuge (unspecified), Bronchitis (unspecified), Aphrodisiac (unspecified), Ache(Loin) (unspecified), Alterative (unspecified), Dyslactation (unspecified), Hepatosis (unspecified), Panacea (unspecified), Parturition (unspecified), Pleurisy (unspecified), Tumor(Glands) (unspecified), Abdominal pain (unspecified), Antineoplastic agents (unspecified), Antirheumatic agents (unspecified), Dyspnea (unspecified), Ear diseases (unspecified), Fractures, bone (unspecified), Gastrointestinal diseases (unspecified), Hemophilia a (unspecified), Hypnotics and sedatives (unspecified), Lactation disorders (unspecified), Liver diseases (unspecified), Neoplasms (unspecified), Pain (unspecified), General tonic for rejuvenation (unspecified), Skin diseases (unspecified), Tuberculosis, pulmonary (unspecified), Ulcer (unspecified), Wounds and injuries (unspecified), Antirheumatic agents (wood), Wounds and injuries (wood)
Human toxicity -
Animal toxicity -

Cultivation

Plants can be grown by seeds or cuttings.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) 22 - 28
Size -
Vigor -
Productivity -

Images

Habit

Alstonia scholaris habit picture by Daniel Barthelemy (cc-by-nc)
Alstonia scholaris habit picture by Dieter Albrecht (cc-by-sa)
Alstonia scholaris habit picture by Islam Asiful (cc-by-sa)

Leaf

Alstonia scholaris leaf picture by Future Foundation (cc-by-sa)
Alstonia scholaris leaf picture by Dieter Albrecht (cc-by-sa)
Alstonia scholaris leaf picture by Lisa Ani (cc-by-sa)

Flower

Alstonia scholaris flower picture by Vineet Kumar Pandey (cc-by-sa)
Alstonia scholaris flower picture by Blue Bottle (cc-by-sa)
Alstonia scholaris flower picture by sanketh kulkarni (cc-by-sa)

Fruit

Alstonia scholaris fruit picture by sanketh kulkarni (cc-by-sa)
Alstonia scholaris fruit picture by Dieter Albrecht (cc-by-sa)
Alstonia scholaris fruit picture by Dieter Albrecht (cc-by-sa)

Distribution

Alstonia scholaris world distribution map, present in Australia, Bangladesh, Brunei Darussalam, Bhutan, China, Micronesia (Federated States of), Indonesia, India, Cambodia, Lao People's Democratic Republic, Sri Lanka, Myanmar, Malaysia, Nepal, Pakistan, Philippines, Papua New Guinea, Solomon Islands, Thailand, Trinidad and Tobago, Taiwan, Province of China, and Viet Nam

Conservation status

Alstonia scholaris threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:60445977-2
WFO ID wfo-0001257712
COL ID C8VD
BDTFX ID -
INPN ID 629055
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Echites pala Alstonia kurzii Echites scholaris Pala scholaris Alstonia scholaris