Herbs or subshrubs, the stems unbranched. Leaves often crowded at the apices of the stems, petiolate, the stipules interpetiolar, large, entire or more often the margin setose pectinate, the processes numerous, unequal. Inflores-cences axillary or terminal, capitate to trichotomously cymose, the bracts entire to multifid, often obscuring-the-flowers. Flowers with the hypanthium oblong or turbinate, the calycine cup short, the lobes' 5, unequal, somewhat foliaceous, persistent; corolla tubular, 5-lobed, the lobes valvate; stamens 5, inserted in the middle of the tube, the anthers-linear, dorsifixed; ovary 2-celled, the ovules nu-merous in each locule, the placentas axile, adnate to the axis. Fruits fleshy, indehiscent, the persistent calyx conspi-cuous.