Plants large, in thick mats or deep wefts, yellow-green to dark green, glossy. Stems irregularly branched. Stem leaves ovate-lanceolate to cordate-ovate; base rounded to subcordate, slightly decurrent; margins revolute, serrate, serrulate, or denticulate; apex acute, acuminate, or abruptly acuminate; costa subpercurrent, with 2 or more shorter, supplementary costae at base; alar cells quadrate to transversely elongate, region extending nearly to costa and an equal distance distally; laminal cells smooth; medial cells uniformly linear-flexuose; apical cells oval-rhombic. Branch leaves similar, narrower, smaller, 2-3 × 1 mm; margins more strongly denticulate; apex narrower, flagelliform, smaller; costa weaker. Specialized asexual reproduction absent. Perigonia on secondary stems or sometimes on side branches, leaves deltoid-apiculate. Perichaetia on primary stems, leaves ecostate. Seta short. Capsule exserted, cylindric, oblong-cylindric, or oblong-ovoid; annulus scant or absent, adherent; operculum oblique-rostrate; exostome teeth subulate, trabeculate or papillose; endostome basal membrane absent, segments weakly developed, narrowly subulate, papillose.
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