Plants medium-sized to large, in loose mats. Stems creeping, sympodial, stoloniferous; secondary stems ascending, irregularly branched, branches few to many, curved or in whorls; paraphyllia absent; radiculose on primary stems. Leaves crowded, imbricate when dry, spreading when moist, ovate, ovate-lanceolate, or cordate-ovate; margins entire, serrate, or serrulate; apex acuminate or acute; ecostate or costa double or more [single], short; alar cells differentiated, many; laminal cells ovate to elliptic, smooth, rarely singly papillose, or occasionally prorate, walls thick; basal cells longer. Sexual condition dioicous; perigonia axillary or borne on secondary stems or branches, deltoid-apiculate or gemmiform; perichaetia borne on main stem or lateral branches. Seta short to elongate. Capsule erect, immersed to exserted, smooth; annulus present or sometimes absent; operculum conic or oblique-rostrate; peristome double; exostome teeth 16, pale; endostome basal membrane low or absent, segments absent, weakly developed, or rarely well developed, cilia absent. Calyptra cucullate, naked or hairy. Spores anisosporous, occasionally large, papillose.
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