Plants medium-sized to large, in rigid tufts, green to brownish, glossy or dull. Stems irregularly branched, usually simple. Stem leaves ovate to ovate-lanceolate; base not cordate, occasionally rounded, not decurrent; margins plane to reflexed or revolute, entire; apex abruptly or gradually slender-acuminate; ecostate; alar cells quadrate, region large; laminal cells smooth or rarely slightly papillose distally over cell lumina; medial laminal cells oblong-rhomboidal to linear; apical cells smooth or papillose, or rarely so abaxially. Branch leaves imbricate, erect-appressed when dry, ascending to spreading when moist, ovate-lanceolate, somewhat concave, larger than stem leaves; margins plane to reflexed, entire or serrulate distally; apex acute to acuminate. Specialized asexual reproduction absent or by small, slender, caducous axillary branchlets. Perigonia axillary, gemmiform. Perichaetia on short, lateral branches, leaves strongly differentiated, greatly elongate, sheathing. Seta short to elongate. Capsule immersed to exserted, ovoid to pyriform; annulus cells in several rows, differentiated; operculum conic or oblique-rostrate; exostome teeth linear, papillose; endostome basal membrane low, segments imperfectly connate, papillose.
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